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Esmeralda buckwheat, Esmeralda wild buckwheat

Thorne's buckwheat, Thorne's wild buckwheat

Habit Herbs, erect to spreading, annual, (0.5–)1–5(–10) dm, glabrous or glandular proximally, mostly grayish. Subshrubs, spreading and matted, not scapose, 0.4–0.8 × 0.4–1(–2.5) dm.
Stems

caudex absent;

aerial flowering stems erect, solid, not fistulose, 0.5–1(–3) dm, glabrous or sparsely glandular proximally.

spreading, with persistent leaf bases, up to 1/4 height of plant;

caudex stems compact;

aerial flowering stems spreading, slender, solid, not fistulose, 0.1–0.2 dm, floccose to slightly tomentose.

Leaves

basal;

petiole 0.5–4 cm, strigose to hispid;

blade obovate to round-ovate, 0.5–2.5 × 0.4–2 cm, sparsely strigose to pilose-hispid, often slightly glandular, grayish, greenish, or reddish on both surfaces, or reddish adaxially, margins plane.

cauline, 1 per node or fasciculate;

petiole 0.01–0.02 cm, tomentose;

blade linear, 0.4–0.6 × 0.05–0.1 cm, densely white-tomentose adaxially, finely villous and green adaxially, margins revolute.

Inflorescences

cymose, open to diffuse, 5–40(–70) × 5–50 cm;

branches not fistulose, glabrous or sparsely glandular near proximal nodes;

bracts 3, scalelike, 0.5–3 × 0.5–2 mm.

umbellate-cymose, compact, 0.5–1 × 0.5–1 cm;

branches usually dichotomous, otherwise with secondaries suppressed, glabrous or nearly so;

bracts 3, scalelike, linear, 0.5–1 mm.

Peduncles

spreading to reflexed, straight, slender to filiform, 0.2–1.5 cm, glabrous.

absent.

Involucres

narrowly turbinate, 0.8–1.8 × 0.5–1.2 mm, glabrous;

teeth 4–5, erect, 0.2–0.4 mm.

1 per node, turbinate, 1.5–2 × 1–1.5 mm, floccose;

teeth 5, erect, 0.4–0.6 mm.

Flowers

1–3 mm;

perianth white to pinkish with greenish or reddish midribs, glabrous, minutely pustulose proximally;

tepals monomorphic, oblanceolate to oblong;

stamens included, 1–1.5(–1.8) mm;

filaments glabrous.

1.5–2 mm;

perianth white, glabrous;

tepals connate proximal 1/4, dimorphic, those of outer whorl obovate, 1–1.5 mm wide, those of inner whorl oblanceolate, 0.8–1 mm wide, connate proximally;

stamens slightly exserted, 2–3 mm;

filaments pilose proximally.

Achenes

light brown, 3-gonous, 1.4–2.5 mm, glabrous.

light brown, 2–2.5 mm, glabrous except for papillate beak.

Eriogonum esmeraldense

Eriogonum thornei

Phenology Flowering May–Jul.
Habitat Copper-rich quartzite gravel on ridges, pinyon woodlands
Elevation 1800 m (5900 ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum thornei is known from a single canyon area in the New York Mountains, San Bernardino County. Shultz elevated Thorne’s wild buckwheat to species status when she proposed E. phoeniceum, here considered a variety of E. microthecum. Eriogonum thornei and E. ericifolium are weakly differentiated, with minimal morphologic differences, and yet they occur on different substrates and are well-removed biogeographically from each other. Thorne’s wild buckwheat is a protected plant in California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowering stems and inflorescence branches glabrous
var. esmeraldense
1. Flowering stems and inflorescence branches glandular to sparsely glandular proximally
var. toiyabense
Source FNA vol. 5, p. 392. FNA vol. 5, p. 255.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. esmeraldense var. esmeraldense, E. esmeraldense var. toiyabense
Synonyms E. ericifolium var. thornei, E. ericifolium subsp. thornei
Name authority S. Watson: Proc. Amer. Acad. Arts 24: 85. (1889) (Reveal & Henrickson) L. M. Shultz: Harvard Pap. Bot. 3: 51. (1998)
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