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spreading buckwheat, spreading wild buckwheat
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Zion buckwheat
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| Shrubs, spreading, not scapose, (1.5–)2–5(–7) × 5–15 dm, grayish- to reddish-brown-tomentose to floccose and gray or, rarely, thinly floccose and greenish. |
Herbs, erect to spreading, 3–8(–10) × 0.5–1.5 dm, glabrous and glaucous, rarely tomentose, greenish. |
spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant; caudex stems absent; aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–0.8 dm, floccose or glabrous. |
spreading to erect, without persistent leaf bases, up to 1/8 height of plant; caudex stems absent; aerial flowering stems erect to spreading, slender to stout, solid or hollow, often fistulose, 1–2.5(–3) dm, glabrous, rarely tomentose. |
cauline, 1 per node; petiole 0.2–0.7 cm, tomentose to floccose; blade oblanceolate to oblong or obovate, (1–)1.5–3 × (0.2–)0.3–0.7 cm, densely white-tomentose abaxially, white-floccose to glabrate or green and glabrous adaxially, margins plane. |
basal, 1 per node; petiole 3–6(–10) cm, tomentose to floccose; blade elliptic or oblong-ovate to ovate, 2–4.5(–6) × 1.5–2.5(–5) cm, lanate to densely tomentose abaxially, thinly floccose or glabrous and green adaxially, margins plane. |
cymose, 10–30(–40) × 10–40 cm; branches dichotomous, white-floccose to glabrate or subglabrous; bracts 3, scalelike, triangular, 0.5–2(–5) mm. |
narrowly virgate or racemose with involucres racemosely disposed throughout or at tips, 20–50(–60) × 5–25 cm, glabrous, rarely tomentose; branches dichotomous, upper secondaries suppressed and bearing (8–)10–15(–20) racemosely arranged involucres; bracts 3, scalelike, triangular, and 2–7 mm, or leaflike, elliptic, and 10–30 × 4–15 mm. |
absent or mostly erect, slender, 0.3–2.5 cm, floccose. |
absent. |
1 per node, turbinate, 1.5–2.5(–3) × 1–2 mm, tomentose to floccose; teeth 5, erect, 0.3–0.6 mm. |
1 per node, turbinate to turbinate-campanulate, 1.5–3 × 1.5–2.5 mm, tomentose; teeth 5, erect, 0.2–0.4 mm. |
2–4 mm; perianth yellow, glabrous; tepals connate proximal 1/4, essentially monomorphic, elliptic to obovate; stamens mostly exserted, 2–4.5 mm; filaments sparsely pilose proximally. |
2–6(–7) mm; perianth white to yellow or bright red, glabrous; tepals connate proximal 1/4 their length, monomorphic, oblong; stamens exserted, 2–5 mm; filaments pilose proximally. |
brown, 2–2.5 mm, glabrous. |
light brown, (3–)4–5(–6) mm, glabrous. |
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| Flowering Jun–Sep. |
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| Sandy to rocky slopes and flats, mixed grassland and sagebrush communities, juniper and montane conifer woodlands |
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| 1200-2500 m (3900-8200 ft) |
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CO; MT; NE; NM; SD; WY
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AZ; UT |
Eriogonum effusum is rather common on the northern Great Plains and along the eastern slope of the Rocky Mountains in central and eastern Colorado, southeastern Montana, western Nebraska, northern New Mexico, southwestern South Dakota, and southeastern Wyoming. Some specimens from Chaffee County, Colorado, are thinly floccose and greenish (Atwood & Welsh 29689, BRY) and thus similar to E. leptocladon. A collection from Pyramid Lake, Washoe County, Nevada (Frandsen & Brown 182, NESH) is clearly mislabeled. A roadside collection of E. effusum gathered near Little America, Sweetwater County, Wyoming, in 1961 (G. Mason 4025, ASU) was an introduction that has not persisted. The spreading wild buckwheat is occasionally merged with Eriogonum microthecum even though the two are morphologically distinct and their ranges do not overlap. Plants in New Mexico are sometimes difficult to distinguish from the related E. leptocladon var. ramosissimum. The species is the food plant for the Rita dotted-blue butterfly (Euphilotes rita). A hybrid between Eriogonum effusum and E. pauciflorum has been named E. ×nebraskense Rydberg [E. multiceps Nees subsp. nebraskense (Rydberg) S. Stokes; E. pauciflorum Pursh var. nebraskense (Rydberg) Reveal]. The hybrid is known from Weld County, Colorado; Cheyenne and Kimball counties, Nebraska; and Converse and Platte counties, Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| FNA vol. 5, p. 248. |
FNA vol. 5, p. 297. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii |
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| E. microthecum var. effusum |
E. racemosum var. zionis |
| Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) |
J. T. Howell: Leafl. W. Bot. 2: 253. (1940) |
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