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spreading buckwheat, spreading wild buckwheat

San Jacinto buckwheat, San Jacinto wild buckwheat

Habit Shrubs, spreading, not scapose, (1.5–)2–5(–7) × 5–15 dm, grayish- to reddish-brown-tomentose to floccose and gray or, rarely, thinly floccose and greenish. Herbs, erect to spreading, annual, 2–9 dm, glabrous and sparsely glandular especially (or at least) near nodes, grayish to greenish.
Stems

spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent;

aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–0.8 dm, floccose or glabrous.

caudex absent;

aerial flowering stems erect, solid, not fistulose, 0.5–1.5 dm, glabrous, glandular distally or sparsely so throughout.

Leaves

cauline, 1 per node;

petiole 0.2–0.7 cm, tomentose to floccose;

blade oblanceolate to oblong or obovate, (1–)1.5–3 × (0.2–)0.3–0.7 cm, densely white-tomentose abaxially, white-floccose to glabrate or green and glabrous adaxially, margins plane.

basal;

petiole 1–4 cm, pilose, slightly winged;

blade oblanceolate to obovate, (0.5–)1–4 × 0.5–1.5 cm, sparsely pilose, glandular and greenish on both surfaces, margins entire.

Inflorescences

cymose, 10–30(–40) × 10–40 cm;

branches dichotomous, white-floccose to glabrate or subglabrous;

bracts 3, scalelike, triangular, 0.5–2(–5) mm.

cymose, open, 30–80 × 10–50 cm;

branches not fistulose, glabrous, sparsely glandular at nodes;

bracts 3, scalelike, 1–2 × 1–2 mm.

Peduncles

absent or mostly erect, slender, 0.3–2.5 cm, floccose.

deflexed, straight, filiform, (0.1–)0.2–0.35 cm, sparsely glandular.

Involucres

1 per node, turbinate, 1.5–2.5(–3) × 1–2 mm, tomentose to floccose;

teeth 5, erect, 0.3–0.6 mm.

turbinate, 1.2–1.5 × 1–1.3 mm, glabrous;

teeth 4, erect or nearly so, 0.3–0.7 mm.

Flowers

2–4 mm;

perianth yellow, glabrous;

tepals connate proximal 1/4, essentially monomorphic, elliptic to obovate;

stamens mostly exserted, 2–4.5 mm;

filaments sparsely pilose proximally.

1.5–2.5 mm;

perianth white with reddish brown midribs, becoming reddish in fruit, rarely yellow, puberulent;

tepals monomorphic, oblong-obovate, sometimes with apiculate tip 1–2 mm;

stamens exserted, 1.5–2 mm;

filaments glabrous.

Achenes

brown, 2–2.5 mm, glabrous.

light brown to brown, lenticular, 1.3–1.7(–2.2) mm, glabrous.

2n

= 40.

Eriogonum effusum

Eriogonum apiculatum

Phenology Flowering Jun–Sep. Flowering May–Nov.
Habitat Sandy to rocky slopes and flats, mixed grassland and sagebrush communities, juniper and montane conifer woodlands Sandy granitic flats and slopes, chaparral communities, oak and conifer woodlands
Elevation 1200-2500 m (3900-8200 ft) (200-)700-2700 m ((700-)2300-8900 ft)
Distribution
from FNA
CO; MT; NE; NM; SD; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Eriogonum effusum is rather common on the northern Great Plains and along the eastern slope of the Rocky Mountains in central and eastern Colorado, southeastern Montana, western Nebraska, northern New Mexico, southwestern South Dakota, and southeastern Wyoming. Some specimens from Chaffee County, Colorado, are thinly floccose and greenish (Atwood & Welsh 29689, BRY) and thus similar to E. leptocladon. A collection from Pyramid Lake, Washoe County, Nevada (Frandsen & Brown 182, NESH) is clearly mislabeled. A roadside collection of E. effusum gathered near Little America, Sweetwater County, Wyoming, in 1961 (G. Mason 4025, ASU) was an introduction that has not persisted.

The spreading wild buckwheat is occasionally merged with Eriogonum microthecum even though the two are morphologically distinct and their ranges do not overlap. Plants in New Mexico are sometimes difficult to distinguish from the related E. leptocladon var. ramosissimum. The species is the food plant for the Rita dotted-blue butterfly (Euphilotes rita). A hybrid between Eriogonum effusum and E. pauciflorum has been named E. ×nebraskense Rydberg [E. multiceps Nees subsp. nebraskense (Rydberg) S. Stokes; E. pauciflorum Pursh var. nebraskense (Rydberg) Reveal]. The hybrid is known from Weld County, Colorado; Cheyenne and Kimball counties, Nebraska; and Converse and Platte counties, Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum apiculatum is restricted to the San Jacinto, Santa Rosa, Palomar, and Cuyamaca mountains of Riverside and San Diego counties. Two collections are discounted as to location: Hall 1025 (MIN), purportedly from the San Bernardino Mountains, certainly was labeled erroneously by the Parish brothers, who redistributed the sheet; and Jaeger s.n. (19 May 1940, DS, IDS, UTC, WTU) from “Falcon Flat” in the Little San Bernardino Mountains appears to be another of the small but significant list of Jaeger specimens with highly dubious label data. No such place is known in Joshua Tree National Park. The phase of the species named var. subvirgatum, with nearly sessile involucres, occurs intermixed with the typical expression.

A yellow-flowered population was found at the eastern end of the San Bernardino Mountains, San Bernardino County. Recent attempts to relocate those plants have been unsuccessful, and they remain undescribed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 248. FNA vol. 5, p. 389.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Synonyms E. microthecum var. effusum
Name authority Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) S. Watson: Proc. Amer. Acad. Arts 17: 378. (1882)
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