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Douglas' buckwheat, Douglas' wild buckwheat

Shockley's buckwheat, Shockley's wild buckwheat

Habit Herbs, matted, occasionally polygamodioecious, 0.4–1.5 × 0.5–6 dm, thinly tomentose to glabrate. Herbs, matted, scapose, 0.3–0.5(–0.7) × (0.5–)1–4(–20) dm, floccose to tomentose, greenish or grayish.
Stems

caudex spreading;

aerial flowering stems erect or nearly so, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, nonflowering aerial branches, 0.4–1.2 dm, with a whorl of 4–8 leaflike bracts ca. midlength, similar to leaf blades, 0.3–1.5 × 0.1–0.3 cm.

matted, sometimes only seemingly so, with persistent leaf bases, up to 1/5 height of plant;

caudex stems matted;

aerial flowering stems absent or scapelike, erect or nearly so, slender, solid, not fistulose, (0.05–)0.1–0.3 dm, floccose to tomentose.

Leaves

in basal rosettes;

petiole 0.05–0.5(–1) cm, tomentose;

blade oblanceolate or elliptic to spatulate, 0.4–1.5(–1.9) × 0.1–0.5 cm, lanate on both surfaces, or tomentose abaxially and slightly less so and greenish adaxially, margins entire, plane.

basal, fasciculate in terminal tufts;

petiole 0.2–0.5 cm, tomentose to floccose;

blade oblanceolate to elliptic or spatulate, (0.2–)0.3–0.8(–1.2) × 0.2–0.4(–0.6) cm, tomentose to floccose, margins plane or slightly thickened.

Inflorescences

capitate, 0.8–1.5 cm wide;

branches absent;

bracts absent immediately below involucre.

capitate, 0.8–2 cm;

branches absent;

bracts 3–5, scalelike, linear to linear-lanceolate, 1.5–4 × 0.6–1 mm.

Peduncles

absent.

Involucres

1 per node, turbinate to turbinate-campanulate, 2.5–3.5 × 2–2.5 mm;

teeth 6–14, lobelike, strongly reflexed, 1.5–4(–6) mm.

2–4(–6) per cluster, campanulate, (2–)2.5–5(–6) × 3–6(–7) mm, rigid, tomentose;

teeth 5–10, erect to spreading, (0.5–)1–3 mm.

Flowers

4–9 mm, including 0.7–2 mm stipelike base;

perianth yellow, cream, or ochroleucous to rose-red, sparsely to densely villous abaxially;

tepals monomorphic, obovate;

stamens exserted, 4–6 mm;

filaments pilose proximally.

2.5–4 mm;

perianth white to rose or yellow, densely pilose;

tepals connate proximally, monomorphic, oblong to obovate;

stamens exserted, 2.5–5 mm;

filaments subglabrous or sparsely pilose proximally.

Achenes

light brown, 3–4.5 mm, glabrous except for pubescent beak.

light brown to brown, 2.5–3 mm, tomentose.

Eriogonum douglasii

Eriogonum shockleyi

Phenology Flowering May–Aug.
Habitat Gravelly or clayey (rarely sandy) flats, washes, and slopes, saltbush, blackbrush, and sagebrush communities, pinyon-juniper woodlands
Elevation (800-)1200-2600 m ((2600-)3900-8500 ft)
Distribution
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum shockleyi is widely distributed in northern Arizona, east-central California, western Colorado, southern Idaho, northwestern New Mexico, Nevada, and Utah. On the Colorado Plateau, it has oblanceolate to spatulate leaf blades usually 0.3–1.2 × 0.3–0.6 cm, scapes 1–3 cm, involucres with long (2–3.5 mm) often spreading teeth, and flowers 3–4 mm. These plants have been distinguished as var. longilobum. The typical Great Basin expression has elliptic leaf blades 0.3–0.6 × 0.3–0.5 cm, scapes 0.5–2 cm, involucres with short (0.5–2 mm) erect teeth, and flowers 2.5–4 mm. Low, compact, hummock-like plants in southwestern Idaho with elliptic leaf blades 0.1–0.3(–0.35) × 0.1–0.15 cm, flowering stems absent or up to 0.5 cm, and involucres 2.5–3 mm with teeth 0.8–1 mm have been termed var. packardiae. These morphologic differences have been shown to be genetically insignificant, however (J. F. Smith and T. A. Bateman 2002). Plants on moving sand dunes at the southeast end of Baking Power Flat in Lincoln County, Nevada, can be up to 2 m across. Further studies may indicate that the various geographic expressions deserve taxonomic recognition.

The Great Basin expression is the food plant of the Bernardino dotted-blue butterfly (Euphilotes bernardino). Members of the species are occasionally found in cultivation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades tomentose abaxially, less so adaxially, narrowly oblanceolate to spatulate; flowers 5-9 mm; e Washington, ne Oregon
var. douglasii
1. Leaf blades usually densely lanate on both surfaces, elliptic to spatulate; flowers 4-8 mm; s Oregon, ne California, n Nevada
→ 2
2. Flowers 4-5(-6) mm; s Oregon, ne California, nw Nevada
var. meridionale
2. Flowers 5-8 mm; ne Nevada
var. elkoense
Source FNA vol. 5, p. 363. FNA vol. 5, p. 286.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. douglasii var. douglasii, E. douglasii var. elkoense, E. douglasii var. meridionale
Synonyms E. caespitosum subsp. douglasii E. pulvinatum, E. shockleyi subsp. candidum, E. shockleyi subsp. longilobum, E. shockleyi var. longilobum, E. shockleyi var. packardiae, E. villiflorum var. candidum
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 9. (1856) S. Watson: Proc. Amer. Acad. Arts 18: 194. (1883)
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