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Douglas' buckwheat, Douglas' wild buckwheat

marum-leaf buckwheat, marum-leaf wild buckwheat, mountain buckwheat

Habit Herbs, matted, occasionally polygamodioecious, 0.4–1.5 × 0.5–6 dm, thinly tomentose to glabrate. Herbs, matted, dioecious, (0.1–)0.5–5 × 2–8 dm, floccose or glabrous.
Stems

caudex spreading;

aerial flowering stems erect or nearly so, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, nonflowering aerial branches, 0.4–1.2 dm, with a whorl of 4–8 leaflike bracts ca. midlength, similar to leaf blades, 0.3–1.5 × 0.1–0.3 cm.

caudex spreading;

aerial flowering stems erect, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, non-flowering aerial branches, (0.1–)0.5–4 dm, floccose or glabrous.

Leaves

in basal rosettes;

petiole 0.05–0.5(–1) cm, tomentose;

blade oblanceolate or elliptic to spatulate, 0.4–1.5(–1.9) × 0.1–0.5 cm, lanate on both surfaces, or tomentose abaxially and slightly less so and greenish adaxially, margins entire, plane.

in loose to compact basal rosettes;

petiole 0.3–2 cm, often tomentose;

blade ovate to oval, 0.3–3 × 0.3–1 cm, densely tannish- to brownish-lanate abaxially, glabrate and green to olive green adaxially, margins entire, plane.

Inflorescences

capitate, 0.8–1.5 cm wide;

branches absent;

bracts absent immediately below involucre.

capitate, 0.5–2 cm wide, mature pistillate plants open and elongate-umbellate, 1–5 × 1–7 cm;

branches usually glabrous;

bracts 5–10, leaflike, 0.2–0.8 cm, often absent immediately below involucre.

Involucres

1 per node, turbinate to turbinate-campanulate, 2.5–3.5 × 2–2.5 mm;

teeth 6–14, lobelike, strongly reflexed, 1.5–4(–6) mm.

1 per node but occasionally appearing congested, turbinate and 2–3 × 1.5–2 mm or campanulate and 4–5 × 5–7 mm, sparsely tomentose or glabrous;

teeth 5–6, erect, 0.4–0.7 mm.

Flowers

4–9 mm, including 0.7–2 mm stipelike base;

perianth yellow, cream, or ochroleucous to rose-red, sparsely to densely villous abaxially;

tepals monomorphic, obovate;

stamens exserted, 4–6 mm;

filaments pilose proximally.

stipelike base 0.5–1 mm;

perianth yellow, glabrous; staminate flowers 1.5–3 mm, tepals ovate; pistillate flowers 4–7 mm, tepals oblanceolate, often becoming reddish in fruit;

stamens exserted, 2–3 mm;

filaments pilose proximally.

Achenes

light brown, 3–4.5 mm, glabrous except for pubescent beak.

light brown to brown, 3.5–5 mm, glabrous except for sparsely pubescent beak.

2n

= 32.

Eriogonum douglasii

Eriogonum marifolium

Distribution
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Eriogonum marifolium is dioecious, with the two sexes having such markedly different morphology that collectors often consider each to represent a different species. In Oregon, male plants of E. marifolium and E. umbellatum var. haussknechtii can be difficult to distinguish. In El Dorado and Placer counties, California, herbarium specimens of E. marifolium and E. incanum can be difficult to separate, as their distribution ranges overlap there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades tomentose abaxially, less so adaxially, narrowly oblanceolate to spatulate; flowers 5-9 mm; e Washington, ne Oregon
var. douglasii
1. Leaf blades usually densely lanate on both surfaces, elliptic to spatulate; flowers 4-8 mm; s Oregon, ne California, n Nevada
→ 2
2. Flowers 4-5(-6) mm; s Oregon, ne California, nw Nevada
var. meridionale
2. Flowers 5-8 mm; ne Nevada
var. elkoense
1. Involucres turbinate, 2-3 × 1.5-2 mm; ne California, wc Nevada, s Oregon, disjunct to Yakima County, Washington
var. marifolium
1. Involucres campanulate, 4-5 × 5-7 mm; nc California
var. cupulatum
Source FNA vol. 5, p. 363. FNA vol. 5, p. 365.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. douglasii var. douglasii, E. douglasii var. elkoense, E. douglasii var. meridionale
E. marifolium var. cupulatum, E. marifolium var. marifolium
Synonyms E. caespitosum subsp. douglasii E. cupulatum, E. marifolium var. apertum
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 9. (1856) Torrey & A. Gray: Proc. Amer. Acad. Arts 8: 161. (1870)
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