Douglas' buckwheat, Douglas' wild buckwheat
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jointed buckwheat, napkinring, napkinring wild buckwheat
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Herbs, matted, occasionally polygamodioecious, 0.4–1.5 × 0.5–6 dm, thinly tomentose to glabrate. |
Herbs, 6–15 × 0.3–0.9 dm, grayish. |
caudex spreading; aerial flowering stems erect or nearly so, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, nonflowering aerial branches, 0.4–1.2 dm, with a whorl of 4–8 leaflike bracts ca. midlength, similar to leaf blades, 0.3–1.5 × 0.1–0.3 cm. |
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5–12 dm, disarticulating into ringlike segments 3–10(–16) mm. |
in basal rosettes; petiole 0.05–0.5(–1) cm, tomentose; blade oblanceolate or elliptic to spatulate, 0.4–1.5(–1.9) × 0.1–0.5 cm, lanate on both surfaces, or tomentose abaxially and slightly less so and greenish adaxially, margins entire, plane. |
petiole 3–8 cm, pilose; blade oblong-ovate, 2.5–7 × 0.7–2(–3) cm, pilose on both surfaces, margins entire. |
capitate, 0.8–1.5 cm wide; branches absent; bracts absent immediately below involucre. |
1–3 × 2–6 cm; bracts scalelike distally, triangular, and 1–3 mm, semileaflike proximally, oblong, and 3–6 mm. |
1 per node, turbinate to turbinate-campanulate, 2.5–3.5 × 2–2.5 mm; teeth 6–14, lobelike, strongly reflexed, 1.5–4(–6) mm. |
2.5–3.5 × 3–5 mm, pilose; teeth 0.5–2 mm. |
4–9 mm, including 0.7–2 mm stipelike base; perianth yellow, cream, or ochroleucous to rose-red, sparsely to densely villous abaxially; tepals monomorphic, obovate; stamens exserted, 4–6 mm; filaments pilose proximally. |
1.5–3 mm; tepals: those of outer whorl oblanceolate, 0.9–1.2 mm wide, those of inner whorl broadly oblanceolate to fan-shaped, 1–1.5 mm wide; stamens 0.8–1.5 mm. |
light brown, 3–4.5 mm, glabrous except for pubescent beak. |
2–2.5 mm. |
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= 40. |
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Flowering May–Oct. |
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Limestone washes, slopes, and cliffs, saltbush and creosote bush communities, juniper woodlands |
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(600-)800-1600 m ((2000-)2600-5200 ft) |
CA; NV; OR; WA
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CA |
Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Rare and restricted to the Cottonwood, Funeral, Grapevine, and Panamint ranges of Death Valley National Park, Inyo County, Eriogonum intrafractum is one of the more remarkable species of the genus. The tall, simple, glabrous stem is branched only near the top, and then with only two to five inflorescence branches. The small leaves are restricted to the base of the plant. The lower part of the stem, when dried, fragments into napkinring-like segments. The rings have no evident value to the species, as they are not capable of vegetative reproduction. Some parenchyma cells at the fracture points are large enough to be seen with the naked eye, and it is likely that they act as compression points. Much as the intervertebral disks in the vertebral column provide a degree of flexibility, so too may these large parenchyma cells allow the top-heavy flowering stem to move in the wind without breaking. The flowers are numerous, each involucre so filled with more than a hundred flowers that it is torn apart by late anthesis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaf blades tomentose abaxially, less so adaxially, narrowly oblanceolate to spatulate; flowers 5-9 mm; e Washington, ne Oregon | var. douglasii |
1. Leaf blades usually densely lanate on both surfaces, elliptic to spatulate; flowers 4-8 mm; s Oregon, ne California, n Nevada | → 2 |
2. Flowers 4-5(-6) mm; s Oregon, ne California, nw Nevada | var. meridionale |
2. Flowers 5-8 mm; ne Nevada | var. elkoense |
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FNA vol. 5, p. 363. |
FNA vol. 5, p. 328. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oligogonum |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Clastomyelon |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. caespitosum subsp. douglasii |
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Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 9. (1856) |
Coville & C. V. Morton: J. Wash. Acad. Sci. 26: 305. (1936) |
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