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divergent buckwheat, divergent wild buckwheat

bastard-sage, bastard-sage buckwheat, Wright's buckwheat, Wright's wild buckwheat

Habit Herbs, spreading, 1–2(–3) dm, puberulent to short-pilose, greenish to reddish. Shrubs, subshrubs, or herbs, rarely scapose, (1–)1.5–10 × 1–15(–18) dm or, if matted, 0.1–2.5(–3) × 0.5–3(–5) dm, , lanate to thinly tomentose, or glabrous, grayish to greenish or reddish.
Stems

aerial flowering stems decumbent to spreading, 0.3–0.5 dm, puberulent to short-pilose.

spreading to erect, with or without persistent leaf bases, up to 1/2 or more height of plant;

caudex stems absent or spreading, occasionally matted;

aerial flowering stems erect to spreading, stout to slender, solid, not fistulose, (0.1–)0.5–4(–6) dm, tomentose, floccose, or glabrous.

Leaves

basal and cauline;

basal: petiole 1–4(–5) cm, puberulent to short-pilose, blade elliptic-oblong to orbiculate, 1–2(–2.5) × 1–2(–2.5) cm, puberulent to short-pilose and green on both surfaces;

cauline: petiole (0–)0.1–2 cm, puberulent to short-pilose, absent distally, blades opposite, oblanceolate to oblong or elliptic, 0.3–1(–1.5) × 0.2–0.8(–1.2) cm, similar to basal blade.

basal and fasciculate in terminal tufts, or cauline and fasciculate, occasionally 1 per node;

petiole 0.02–0.5(–1) cm, tomentose to floccose;

blade oblanceolate to broadly elliptic, 0.1–3 × 0.1–1 cm, tomentose to floccose, sometimes subglabrous or glabrous and green adaxially, margins plane, sometimes revolute.

Inflorescences

cymose, distally uniparous due to suppression of secondary branches, diffuse, 5–25 × 10–45 cm;

branches puberulent;

bracts 1–3(–5) × 1–2 mm.

virgate or cymose with involucres disposed at tips racemosely arranged involucres, rarely capitate, (1–)5–20 × (1–)10–40 cm;

branches dichotomous, tomentose, floccose, or glabrous;

bracts 3, triangular, scalelike, 0.5–3.5 mm.

Peduncles

absent.

absent.

Involucres

somewhat appressed to branches, campanulate, 1–2 × 1–2 mm, pilose;

teeth 5, lobelike, spreading to somewhat reflexed, 0.7–1.5 mm.

1 per node, turbinate to narrowly campanulate, (0.7–)1–4 × 1–2.5 mm, tomentose, floccose, or glabrous;

teeth 5, erect, 0.3–1 mm.

Flowers

(1–)1.5–2 mm;

perianth yellow, rarely pale yellow, hispidulous and glandular with yellowish-white hairs;

tepals monomorphic, oblong-lanceolate to oblong-ovate;

stamens included, 0.7–1.5 mm;

filaments pilose proximally.

1–4 mm;

perianth white to pink or rose, glabrous;

tepals connate proximal 1/4, monomorphic, obovate;

stamens exserted, 1.5–4 mm;

filaments glabrous or sparsely pilose proximally.

Achenes

light brown, 3-gonous, 1.5–2 mm.

light brown to brown, (1–)1.5–3 mm, glabrous.

Eriogonum divaricatum

Eriogonum wrightii

Phenology Flowering May–Oct.
Habitat Heavy clay flats and slopes, saltbush, greasewood, and sagebrush communities, pinyon-juniper woodlands
Elevation 1100-2300(-2500) m (3600-7500(-8200) ft)
Distribution
from FNA
AZ; CO; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; NV; TX; UT; including nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Eriogonum divaricatum basically is a species of the Colorado Plateau of southwestern Wyoming (Sublette, Sweetwater, and Uinta counties), eastern Utah (Carbon, Emery, Garfield, Grand, Kane, San Juan, Sevier, and Wayne counties), western Colorado (Mesa and Montezuma counties), northern Arizona (Apache, Coconino, and Navajo counties), and northwestern New Mexico (McKinley and San Juan counties), where it is infrequent to common but only rarely abundant. There are several disjunct populations well removed from that core area; the species occurs in Beaver and Millard counties of west-central Utah, and in Clayhole Valley, Mohave County, Arizona. In New Mexico, it was found around San Ysidro, Sandoval County, in 1926. The most extreme disjunctions were two populations found in Argentina during the growing season of 1899–1900. Those plants were named Eriogonum ameghinoi Spegazzini, which ultimately became the only species of the genus Sanmartinia M. Buchinger. Probably introduced there by migrating birds, the species did not persist in South America (J. L. Reveal 1981b).

The divergent wild buckwheat was used ceremonially by the Navajo (Diné) people in one of their snake dances, and portions of the plant were smoked in the treatment of snakebite (A. Clifford, pers. comm.; L. C. Wyman and S. K. Harris 1951).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 9 (6 in the flora).

Eriogonum wrightii is subdivided into several varieties most of which are distinct, although a few have rather indistinct boundaries. Some of the variation has yet to be fully resolved, especially in the var. nodosum complex where, at least in Mexico, one additional expression remains to be named. A clear distinction between var. subscaposum and E. kennedyi is not possible in southern California (see discussion below). Nearly all of the varieties of E. wrightii are in cultivation, although the most elegant (var. olanchense) has yet to be so honored.

Few ethnobotanical uses of bastard-sage are reported in the literature. L. C. Wyman and S. K. Harris (1951) noted that the Kayenta Navajo use it (var. wrightii) as an emetic, while M. L. Zigmond (1981) stated that the Kawaiisu used the pounded seeds (probably of var. subscaposum) in a beverage or as a dry meal. Members of the species are food plants for the rare Rita dotted-blue butterfly (Euphilotes rita), the Pacific dotted-blue (E. enoptes), the veined blue (Plebeius neurona), and the Mormon metalmark (Apodemia mormo mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants loosely to compactly matted or herbs
→ 2
1. Plants shrubs or subshrubs
→ 3
2. Plants 0.5-2.5(-3) dm; leaf blades 0.5-1(-1.2) cm; involucres 1.5-4 mm; e and s California,wc Nevada
var. subscaposum
2. Plants 0.1-0.3(-0.6) dm; leaf blades 0.1-0.25 cm; involucres 0.8-1.7(-2) mm; se California
var. olanchense
3. Flowering stems and branches grayish, lanate to densely tomentose; sw Arizona, se California
var. nodosum
3. Flowering stems and inflorescence branches whitish, reddish, or greenish, tomentose to floccose; widespread
→ 4
4. Petiole bases forming distinct ring around stem; leaf blades 0.2-0.6(-1) × 0.1-0.3(-0.4) cm; s California
var. membranaceum
4. Petiole bases not forming distinct ring around stem; leaf blades 0.5-3 × 0.2-1 cm; sw United States, Mexico
→ 5
5. Leaf blades 0.5-1.5 × 0.2-0.5(-0.7) cm; involucres 2-2.5 mm; flowers 2.5-3.5 mm; se California to w Texas
var. wrightii
5. Leaf blades 1.5-3 × 0.5-1 cm; involucres 3-4 mm; flowers 3-4 mm; c and nw California
var. trachygonum
Source FNA vol. 5, p. 429. FNA vol. 5, p. 289.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oregonium Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. zionis
Subordinate taxa
E. wrightii var. membranaceum, E. wrightii var. nodosum, E. wrightii var. olanchense, E. wrightii var. subscaposum, E. wrightii var. trachygonum, E. wrightii var. wrightii
Synonyms E. trachygonum subsp. wrightii
Name authority Hooker: Hooker’s J. Bot. Kew Gard. Misc. 5: 265. (1853) Torrey ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 15. (1856)
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