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divergent buckwheat, divergent wild buckwheat
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Heerman buckwheat, Heermann's buckwheat, Heermann's wild buckwheat
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| Herbs, spreading, 1–2(–3) dm, puberulent to short-pilose, greenish to reddish. |
Shrubs and subshrubs, spreading to rounded and occasionally erect, not scapose, (0.5–)1–20 × 2–25 dm, glabrous or occasionally floccose, sometimes scabrellous, greenish, infrequently grayish. |
aerial flowering stems decumbent to spreading, 0.3–0.5 dm, puberulent to short-pilose. |
spreading or erect, without persistent leaf bases, up to 1/2 height of plant; caudex stems absent; aerial flowering stems erect or nearly so, slender to stout, solid, not fistulose, 0.02–0.5 dm, thinly tomentose or glabrous. |
basal and cauline; basal: petiole 1–4(–5) cm, puberulent to short-pilose, blade elliptic-oblong to orbiculate, 1–2(–2.5) × 1–2(–2.5) cm, puberulent to short-pilose and green on both surfaces; cauline: petiole (0–)0.1–2 cm, puberulent to short-pilose, absent distally, blades opposite, oblanceolate to oblong or elliptic, 0.3–1(–1.5) × 0.2–0.8(–1.2) cm, similar to basal blade. |
cauline, 1 per node, quickly deciduous; petiole 0.1–1.5 cm, floccose or glabrous; blade linear, oblanceolate or spatulate or elliptic, or oblong, (0.4–)1–2(–4) × 0.1–0.8 cm, tomentose to floccose or glabrous abaxially, floccose to thinly floccose or glabrous adaxially, margins plane. |
cymose, distally uniparous due to suppression of secondary branches, diffuse, 5–25 × 10–45 cm; branches puberulent; bracts 1–3(–5) × 1–2 mm. |
cymose or racemose, 1–25(–30) × 1–30(–35) cm; branches dichotomous, sometimes with secondaries suppressed, smooth or angled to ridged and grooved, glabrous or occasionally floccose or scabrous; bracts 3, scalelike, 0.3–2 mm. |
absent. |
absent. |
somewhat appressed to branches, campanulate, 1–2 × 1–2 mm, pilose; teeth 5, lobelike, spreading to somewhat reflexed, 0.7–1.5 mm. |
1 per node, narrowly turbinate or campanulate, 0.7–3 × 0.7–4 mm, glabrous, infrequently floccose; teeth 5, erect, 0.3–0.7 mm. |
(1–)1.5–2 mm; perianth yellow, rarely pale yellow, hispidulous and glandular with yellowish-white hairs; tepals monomorphic, oblong-lanceolate to oblong-ovate; stamens included, 0.7–1.5 mm; filaments pilose proximally. |
(1.5–)2–4 mm; perianth white, yellowish white, pink, or reddish, glabrous; tepals connate proximal 1/4, dimorphic, those of outer whorl obovate to orbiculate, those of inner whorl narrowly lanceolate to oblong; stamens exserted, 2–5 mm; filaments pilose proximally. |
light brown, 3-gonous, 1.5–2 mm. |
light brown to brown, 2–5 mm, glabrous. |
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| Flowering May–Oct. |
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| Heavy clay flats and slopes, saltbush, greasewood, and sagebrush communities, pinyon-juniper woodlands |
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| 1100-2300(-2500) m (3600-7500(-8200) ft) |
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AZ; CO; NM; UT; WY
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AZ; CA; NV; UT
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Eriogonum divaricatum basically is a species of the Colorado Plateau of southwestern Wyoming (Sublette, Sweetwater, and Uinta counties), eastern Utah (Carbon, Emery, Garfield, Grand, Kane, San Juan, Sevier, and Wayne counties), western Colorado (Mesa and Montezuma counties), northern Arizona (Apache, Coconino, and Navajo counties), and northwestern New Mexico (McKinley and San Juan counties), where it is infrequent to common but only rarely abundant. There are several disjunct populations well removed from that core area; the species occurs in Beaver and Millard counties of west-central Utah, and in Clayhole Valley, Mohave County, Arizona. In New Mexico, it was found around San Ysidro, Sandoval County, in 1926. The most extreme disjunctions were two populations found in Argentina during the growing season of 1899–1900. Those plants were named Eriogonum ameghinoi Spegazzini, which ultimately became the only species of the genus Sanmartinia M. Buchinger. Probably introduced there by migrating birds, the species did not persist in South America (J. L. Reveal 1981b). The divergent wild buckwheat was used ceremonially by the Navajo (Diné) people in one of their snake dances, and portions of the plant were smoked in the treatment of snakebite (A. Clifford, pers. comm.; L. C. Wyman and S. K. Harris 1951). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 8 (8 in the flora). As Eriogonum heermannii is here circumscribed, the number of varieties is dramatically decreased from past presentations, with E. apachense reduced to synonymy under a now greatly expanded var. argense. Also included in that variety is the more stoutly branched var. subracemosum. The southern var. heermannii of basically desert ranges and the more northern var. occidentale of the Coast Ranges in California are maintained, but their separation is more traditional than certain. The fragile and bulky nature of many dried, often poorly prepared specimens, and the tendency for leaves to fall away have made varietal identification within E. heermannii difficult. Eriogonum heermannii varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi) and the Mormon metalmark (Apodemia mormo mormo). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Flowering stems and inflorescence branches round or angled, scabrellous or papillate-scabrous; plants often densely branched | → 2 |
1. Flowering stems and inflorescence branches round, not angled, usually smooth, glabrous or thinly tomentose; plants sparsely branched | → 3 |
2. Flowering stems and inflorescence branches round, usually distinctly scabrellous or infrequently papillate-scabrous; Arizona, se California, Nevada | var. argense |
2. Flowering stems and inflorescence branches sharply ridged and deeply grooved, minutely scabrellous; nw Arizona, se California, s Nevada, sw Utah | var. sulcatum |
3. Involucres not racemosely arranged on inflorescence branches, or only last 2-3 so disposed; inflorescences diffusely branched, glabrous; Great Basin or northeast edge of Mojave Desert | → 4 |
3. Involucres racemosely arranged along inflorescence branches or at least at tips of branches; inflorescences openly branched, glabrous or floccose to thinly tomentose; Mojave and Sonoran deserts | → 5 |
4. Inflorescence branches mostly slender, smooth, not ridged, not spinose; inflorescences 3-15(-23) × 5-20 cm; subshrubs or shrubs, 3-7 × 5-12(-15) dm; Great Basin, ec California and Nevada | var. humilius |
4. Inflorescence branches stoutish, faintly ridged and grooved, spinose; inflorescences 3-7(-10) × 3-10(-12) cm; shrubs, 1-3 × 1.5-5(-8) dm; Mojave Desert, sw Utah and nw Arizona | var. subspinosum |
5. Inflorescence branches thinly tomentose to floccose; s California, s Nevada, nw Arizona | var. floccosum |
5. Inflorescence branches glabrous; s Nevada, California | → 6 |
6. Involucres racemosely arranged; inflorescence branches whiplike; s Nevada | var. clokeyi |
6. Involucres racemosely arranged only distally; inflorescence branches not whiplike; California | → 7 |
7. Leaf blades 0.5-1.5 cm, glabrous abaxially; inflorescence branches stout; sc California | var. heermannii |
7. Leaf blades 1.5-3(-4) cm, mostly tomentose to floccose abaxially (at least in early anthesis); inflorescence branches slender; sw California | var. occidentale |
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| FNA vol. 5, p. 429. |
FNA vol. 5, p. 304. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Oregonium |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. geniculatum |
| Hooker: Hooker’s J. Bot. Kew Gard. Misc. 5: 265. (1853) |
Durand & Hilgard: in War Department [U.S.], Pacif. Railr. Rep. 5(3): 14. (1857) |
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