corymbed wild buckwheat, crispleaf buckwheat, crispleaf wild buckwheat, white wild buckwheat
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San Jacinto buckwheat, San Jacinto wild buckwheat
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Shrubs or subshrubs, spreading, rounded, occasionally erect, rarely somewhat matted, not scapose, (0.5–)1.5–8(–15) × (2–)3–15(–23) dm, grayish- to reddish-brown-tomentose to floccose or glabrous, grayish or greenish. |
Herbs, erect to spreading, annual, 2–9 dm, glabrous and sparsely glandular especially (or at least) near nodes, grayish to greenish. |
spreading or erect, often with persistent leaf bases, up to 3/4 or more height of plant; caudex stems absent or somewhat matted; aerial flowering stems erect or nearly so, slender or occasionally stout, solid, not fistulose, (0.1–)1–2 dm, tomentose to floccose, occasionally glabrous. |
caudex absent; aerial flowering stems erect, solid, not fistulose, 0.5–1.5 dm, glabrous, glandular distally or sparsely so throughout. |
cauline, 1 per node; petiole 0.1–1.5 cm, tomentose to floccose; blade lanceolate to oblanceolate or elliptic to nearly orbiculate, rarely cordate, (0.5–)1–3(–4.5) × (0.3–)0.5–3(–3.5) cm, densely white-, tannish- or brownish-tomentose on both surfaces or less so to nearly glabrous and green adaxially, margins occasionally crenulate. |
basal; petiole 1–4 cm, pilose, slightly winged; blade oblanceolate to obovate, (0.5–)1–4 × 0.5–1.5 cm, sparsely pilose, glandular and greenish on both surfaces, margins entire. |
cymose, rarely capitate or umbellate, diffuse to rather open, (1–)3–20 × 2–25(–30) cm; branches dichotomous, tomentose, floccose, or rarely glabrous; bracts 3, scalelike, usually triangular, and 1–3(–6) mm, or leaflike, 10–25 mm, and similar to leaf blades. |
cymose, open, 30–80 × 10–50 cm; branches not fistulose, glabrous, sparsely glandular at nodes; bracts 3, scalelike, 1–2 × 1–2 mm. |
absent. |
deflexed, straight, filiform, (0.1–)0.2–0.35 cm, sparsely glandular. |
1 per node, turbinate, (1–)1.5–3.5 × 1–2(–2.5) mm; teeth 5, erect, 0.3–1 mm. |
turbinate, 1.2–1.5 × 1–1.3 mm, glabrous; teeth 4, erect or nearly so, 0.3–0.7 mm. |
(1.5–)2–3.5 mm; perianth white to cream, pink, or pale yellow to yellow, glabrous or rarely sparsely pilose; tepals connate proximal 1/4–1/3, essentially monomorphic, oblanceolate to spatulate; stamens included to slightly exserted, 1–4(–5) mm; filaments typically pilose proximally. |
1.5–2.5 mm; perianth white with reddish brown midribs, becoming reddish in fruit, rarely yellow, puberulent; tepals monomorphic, oblong-obovate, sometimes with apiculate tip 1–2 mm; stamens exserted, 1.5–2 mm; filaments glabrous. |
brown, 2–2.5(–3) mm, glabrous except for occasional papillate beak. |
light brown to brown, lenticular, 1.3–1.7(–2.2) mm, glabrous. |
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= 40. |
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Flowering May–Nov. |
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Sandy granitic flats and slopes, chaparral communities, oak and conifer woodlands |
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(200-)700-2700 m ((700-)2300-8900 ft) |
AZ; CO; NM; NV; TX; UT; WY
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CA
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Varieties 8 (8 in the flora). Eriogonum corymbosum is a difficult complex of overlapping expressions, some of which are maintained here as taxonomically significant. Although perianth color is used to group the varieties, this feature is not consistent even in single populations. Therefore, population trends in perianth color must be noted in the field. Most of the varieties are then distinguished on the basis of leaf characters, and again, considerable variation can be seen in some populations. Still, the combination of flower color, leaf features, and geographic distribution should prove useful in distinguishing the varieties. S. L. Welsh et al. (2003) alluded to hybrid combinations involving Eriogonum corymbosum and other species. Aside from the instances involving E. brevicaule, discussed below, none has been confirmed. Most of the putative hybrids are misidentified specimens of E. lonchophyllum or collections of var. corymbosum in which the leaf-margins are not decidedly crisped, a feature usually seen only in fully mature plants. Eriogonum corymbosum was widely used by Native Americans. P. A. Vestal (1940) reported that the Hopi pressed boiled stalks into cakes that, when dried, were eaten with salt. J. W. Fewkes (1896) indicated that boiled leaves were mixed with cornmeal and water, and then baked into a kind of bread. S. A. Weber and P. D. Seaman (1985) indicated that A. F. Whiting was aware of a decoction of leaves (probably from var. glutinosum) being used for headaches. Variety glutinosum also was used primarily to treat tuberculosis, or at least as a cough medicine (D. E. Moerman 1986). Some of the expressions of Eriogonum corymbosum are attracting the interest of gardeners, a few are coming into cultivation, and several selections are now being developed. The plants are slow growing but can be transplanted with some degree of success. Members of the varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eriogonum apiculatum is restricted to the San Jacinto, Santa Rosa, Palomar, and Cuyamaca mountains of Riverside and San Diego counties. Two collections are discounted as to location: Hall 1025 (MIN), purportedly from the San Bernardino Mountains, certainly was labeled erroneously by the Parish brothers, who redistributed the sheet; and Jaeger s.n. (19 May 1940, DS, IDS, UTC, WTU) from “Falcon Flat” in the Little San Bernardino Mountains appears to be another of the small but significant list of Jaeger specimens with highly dubious label data. No such place is known in Joshua Tree National Park. The phase of the species named var. subvirgatum, with nearly sessile involucres, occurs intermixed with the typical expression. A yellow-flowered population was found at the eastern end of the San Bernardino Mountains, San Bernardino County. Recent attempts to relocate those plants have been unsuccessful, and they remain undescribed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Perianths usually pale yellow to yellow, occasionally white; s Utah, n Arizona, se Nevada | → 2 |
1. Perianths white to cream or pink; sw Wyoming s through e Utah and w Colorado to nw New Mexico and Arizona | → 4 |
2. Inflorescence branches glabrous or nearly so; sw Utah | var. aureum |
2. Inflorescence branches tomentose to floccose | → 3 |
3. Leaf blades greenish-floccose adaxially; Colorado Plateau, n Arizona, s Utah | var. glutinosum |
3. Leaf blades silvery-floccose adaxially; Mojave Desert, se Nevada, sc Utah | var. nilesii |
4. Leaf blades elliptic-oblong to nearly orbiculate, 1-3(-3.5) × 1-2.5(-3.5) cm | → 5 |
4. Leaf blades lanceolate to oblanceolate or elliptic, (0.5-)1-3(-4.5) × (0.3-)0.5-1.5 cm | → 6 |
5. Leaf blades elliptic-oblong to oblong or ovate, rarely cordate, densely white- to tan-lanate or less so to brownish-floccose adaxially; plants brownish-white- tomentose; flowers 2-2.5(-3) mm | var. velutinum |
5. Leaf blades usually broadly ovate to nearly orbiculate, densely grayish-tomentose abaxially, sometimes tomentose to floccose on both surfaces, or more often subglabrous or glabrous and green adaxially; plants greenish-tomentose; flowers 2.5-3 mm | var. orbiculatum |
6. Plants sprawling subshrubs, 0.5-1.5 dm, occasionally shrubs, 2-4.5 dm; leaf blades 0.5-2 × 0.3-0.6(-0.8) cm; subalpine in bristlecone pine communities | var. heilii |
6. Plants erect subshrubs or shrubs, 3-10 dm; leaf blades 1-3(-4.5) × (0.3-)0.5-1.5 cm; desert scrub to montane forest communities | → 7 |
7. Leaves cauline 2 or more length of flowering stems; n Arizona, w Colorado, e and s Utah, sw Wyoming | var. corymbosum |
7. Leaves cauline 4 or less length of flowering stems; c and e Utah | var. revealianum |
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FNA vol. 5, p. 255. |
FNA vol. 5, p. 389. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. effusum subsp. corymbosum |
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Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 17. (1856) |
S. Watson: Proc. Amer. Acad. Arts 17: 378. (1882) |
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