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shortstem buckwheat, shortstem wild buckwheat

drop-leaf wild buckwheat, dropleaf buckwheat

Habit Herbs, matted, cespitose, pulvinate, erect or spreading, sometimes scapose, (0.3–)1–5 × 1–5(–8) dm, tomentose to floccose or glabrous, grayish or greenish to green. Herbs, matted, not scapose, 0.3–1 × 1–2 dm, glabrous, green.
Stems

matted to spreading, occasionally with persistent leaf bases, up to 1/4 or more height of plant;

caudex stems matted or spreading;

aerial flowering stems spreading to erect or nearly so, slender, rarely stout, solid, not fistulose, (0.4–)0.5–2(–2.5) dm, glabrous, floccose, or sparsely to densely tomentose to lanate.

spreading, usually with persistent leaf bases, up to 1/5 height of plant;

caudex stems matted;

aerial flowering stems mostly erect, slender, solid, not fistulose, 0.03–0.1 dm, glabrous or floccose or sparsely tomentose.

Leaves

basal or more commonly sheathing 1–7(–15) cm up stem, 1 per node;

petiole 0.2–2(–4) cm, tomentose to floccose;

blade linear, oblanceolate, or spatulate to elliptic, (0.2–)1–10(–12) × 0.1–0.9(–1.2) cm, densely tomentose abaxially, less so to floccoseadaxially, margins plane or revolute, sometimes crenulate.

basal, 1 per node but congested;

petiole 0.5–1 cm, tomentose to floccose or glabrous;

blade linear to linear-oblanceolate, (2–)3–6 × 0.1–0.3 cm, densely white-tomentose abaxially, less so or glabrous and green adaxially, margins revolute.

Inflorescences

cymose, subumbellate, umbellate, or capitate, (1–)3–10(–25) × (0.7–)1–10(–15) cm;

branches dichotomous, sometimes absent, tomentose to floccose or glabrous;

bracts 3, triangular, scalelike, 1–3(–5) mm.

cymose-umbellate, 1.5–3 × 1.5–3 cm;

branches dichotomous, glabrous;

bracts 3, semileaflike, usually triangular, 3–5 mm.

Peduncles

absent or erect, 0.3–3 cm, tomentose to floccose or glabrous.

stout, 0.1–0.4 cm, glabrous.

Involucres

1 per node or 3–7(–9) per cluster, turbinate to turbinate-campanulate, 1.5–4(–5) × (1–)1.5–3(–3.5) mm, tomentose to floccose or glabrous;

teeth 5, erect to spreading, 0.3–1 mm.

1 per node, turbinate-campanulate, 2.5–3.5(–4.5) × 2–3 mm, glabrous except for cottony tomentum between teeth;

teeth 5, erect, 0.3–0.5 mm.

Flowers

(1–)2–4 mm;

perianth various shades of white to cream or yellow, glabrous or pubescent;

tepals connate proximal 1/4–1/3, monomorphic, lanceolate, oblong to obovate or ovate to oval;

stamens exserted, 2–4 mm;

filaments pilose basally.

2–3.5 mm;

perianth white, glabrous;

tepals connate proximal 1/4–1/3, monomorphic, oblanceolate to elliptic;

stamens exserted, 3–4 mm;

filaments sparsely pilose proximally.

Achenes

light brown to brown, 2–3 mm, glabrous except for roughened to papillate beak.

brown, 2.5–3.5 mm, glabrous.

Eriogonum brevicaule

Eriogonum exilifolium

Phenology Flowering Jun–Sep.
Habitat Clay hills and flats or granitic sandy slopes, mixed grassland and sagebrush communities
Elevation 2200-2600 m (7200-8500 ft)
Distribution
from FNA
CO; ID; MT; NE; NV; SD; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; WY
[BONAP county map]
Discussion

Varieties 8 (8 in the flora).

Eriogonum brevicaule is highly variable, and the variation has yet to be fully resolved taxonomically. The expressions recognized here will encompass the vast majority of populations. The extreme variation previously under the name var. laxifolium is now reduced with the recognition of var. bannockense (low-elevation or northern phase), var. nanum, and var. caelitum (high-elevation, southern phases).

Essentially all of the following species (28–63 below) belong to the Eriogonum brevicaule complex. Eriogonum desertorum, E. loganum, E. spathulatum, E. ostlundii, and E. artificis are allied to the complex associated with var. laxifolium, while E. natum is related to var. cottamii. Eriogonum viridulum and E. ephedroides are allied to E. brevicaule var. brevicaule as are E. contortum and E. acaule. Eriogonum brandegeei is also related, but exactly how is less certain. Allied to this complex of species are on the one hand those related to E. batemanii, and on the other all of the matted perennials belonging to the E. ochrocephalum complex. Essentially all of these species form relatively small populations on discrete edaphic sites and are well isolated one from the other. Unfortunately, a clear separation of E. brevicaule from E. desertorum, E. loganum, and E. spathulatum is not always possible.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum exilifolium is known only from Grand, Jackson, and Larimer counties, Colorado, and from Carbon and Albany counties, Wyoming. The drop-leaf wild buckwheat is a “species of concern” in Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowering stems and inflorescence branches glabrous
→ 2
1. Flowering stems and inflorescence branches floccose to tomentose or lanate
→ 3
2. Plants not pulvinate or cespitose, (0.8-)1.5-5 dm; inflorescences cymose, open, divided 3 times or more; widespread
var. brevicaule
2. Plants pulvinate and cespitose, 0.3-1.5(-1.8) dm; inflorescences capitate or umbellate to cymose and divided 1-2 times; Utah
var. nanum
3. Perianths pubescent; se Montana, ne Wyoming
var. canum
3. Perianths glabrous; sc Idaho, ne Nevada, Utah, sw Wyoming, not se Montana or ne Wyoming
→ 4
4. Inflorescences divided (2-)3-5 times; perianths usually yellow, rarely ochroleucous
→ 5
4. Inflorescences capitate or umbellate to cymose and divided 1-2 times; perianths ochroleucous or yellow
→ 6
5. Flowers (1-)1.5-2.5 mm; sw Wyoming
var. micranthum
5. Flowers (2.5-)3-4 mm; Utah
var. cottamii
6. Leaf blades linear to narrowly oblanceolate; inflorescences capitate or divided
→ 7
6. Leaf blades linear, oblanceolate, or narrowly oblanceolate to elliptic; inflorescences capitate. [8. Shifted to left margin.—Ed.]
→ 8
7. Leaf blades (1.5-)3-9(-12) × 0.1-0.5(-0.7) cm, tomentose abaxially, less so and grayish or occasionally greenish adaxially, margins usually revolute, occasionally plane; perianths ochroleucous or yellow; n Utah, se Idaho, below 2800 m
var. laxifolium
7. Leaf blades 0.2-4.5(-5) × (0.2-)0.3-0.6(-0.7) cm, tomentose abaxially, thinly floccose and bright green adaxially, margins plane or slightly thickened; perianths yellow; c Utah, above 2700 m
var. caelitum
8. Leaf blades and flowering stems bright green under the thinly floccose pubescence; ne Nevada and nw Utah
var. laxifolium
8. Leaf blades and flowering stems usually grayish to dull greenish under tomentum, rarely thinly floccose; se Idaho and sw Wyoming s to n Utah and ne Nevada
→ 9
9. Leaf margins crenulate; leaf blades narrowly oblanceolate to narrowly elliptic, (0.3-)0.5-1.5(-2) × 0.2-0.5(-7) cm, densely white-tomentose abaxially, floccose and greenish adaxially
var. nanum
9. Leaf margins usually plane (rarely crenulate in Wyoming); leaf blades narrowly oblanceolate to oblanceolate, (0.8-)1-4(-4.5) × (0.3-)0.4-0.8 cm, densely tomentose abaxially, tomentose to floccose and grayish to greenish adaxially
var. bannockense
Source FNA vol. 5, p. 264. FNA vol. 5, p. 263.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. brevicaule var. bannockense, E. brevicaule var. brevicaule, E. brevicaule var. caelitum, E. brevicaule var. canum, E. brevicaule var. cottamii, E. brevicaule var. laxifolium, E. brevicaule var. micranthum, E. brevicaule var. nanum
Synonyms E. campanulatum subsp. brevicaule
Name authority Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) Reveal: Great Basin Naturalist 27: 114, fig. 1. (1967)
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