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island buckwheat, Santa Cruz Island buckwheat, Santa Cruz Island wild buckwheat

fine branch wild buckwheat, sand buckwheat, sand wild buckwheat

Habit Shrubs, erect to spreading, 6–15(–20) × 5–30 dm, glabrous and cinereous, reddish. Shrubs, erect to spreading, not scapose, (2–)3–10(–12) × 5–15(–20) dm, white-tomentose to floccose, or green and nearly glabrous.
Stems

spreading to erect, occasionally with persistent leaf bases, up to 3/4 or more height of plant;

caudex stems absent;

aerial flowering stems erect, stout, solid, not fistulose, 0.5–1(–1.5) dm, glabrous.

spreading, often without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent or spreading in moving sand;

aerial flowering stems spreading, slender, solid, not fistulose, 0.3–1 dm, white-tomentose, floccose to glabrate or glabrous.

Leaves

cauline, 1 per node;

petiole 0.1–0.5 cm, tomentose;

blade linear to narrowly oblong, 2–4(–5) × 0.1–0.4(–0.6) cm, white-tomentose abaxially, cinereous to glabrate and greenish adaxially, margins often revolute.

cauline, 1 per node;

petiole 0.2–0.5 cm, tomentose to floccose;

blade linear-lanceolate to linear-oblanceolate to narrowly oblong, or lanceolate to narrowly elliptic, 1.5–4 × 0.2–0.8(–1.2) cm, densely white-tomentose abaxially, less so and greenish adaxially, margins plane, infrequently revolute.

Inflorescences

cymose, 5–10 × 5–15(–20) cm;

branches dichotomous, glabrous;

bracts 1–3, leaflike, lanceolate to oblong, 0.5–2 × 0.3–1.5 cm.

cymose, open, 5–40 × 10–50 cm;

branches dichtomous proximally, often with involucres racemosely arranged proximally, tomentose to floccose or glabrate, occasionally glabrous;

bracts 3, scalelike, linear to triangular, 1–3(–6) mm.

Peduncles

absent or erect, stout, 0.1–0.5 cm, cinereous.

absent.

Involucres

1 per node, campanulate, 2–3 × 2.5–4 mm, cinereous;

teeth 5–7, erect, 0.5–1.5 mm.

1 per node, turbinate to turbinate-campanulate, 1.5–3 × 1–2 mm, tomentose to floccose or glabrous;

teeth 5, erect, 0.4–0.7 mm.

Flowers

2–3.5(–4) mm;

perianth white to pinkish, villous abaxially;

tepals connate proximally, monomorphic, oblanceolate to narrowly obovate;

stamens exserted, 3–5 mm;

filaments glabrous.

(2–)2.5–3.5 mm;

perianth white or pale yellow to yellow, glabrous;

tepals connate proximal 1/4–1/3, essentially monomorphic, oblong to broadly obovate;

stamens slightly exserted, 2–4 mm;

filaments sparsely pilose proximally.

Achenes

brown, 2.5–3.5 mm, glabrous.

light brown, 2.5–3.5 mm, glabrous except for minutely papillate beak.

2n

= 40.

Eriogonum arborescens

Eriogonum leptocladon

Phenology Flowering Apr–Oct.
Habitat Rocky slopes and canyon walls, coastal scrub communities
Elevation 10-600 m (0-2000 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Eriogonum arborescens is local and occasionally rare on Santa Cruz, Santa Rosa, and Anacapa islands in Santa Cruz and Ventura counties. The species is cultivated, and populations have become naturalized on the mainland from San Mateo County south to San Diego, in large part because of planting along highways. Every effort should be made to remove these naturalized populations from the mainland.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Eriogonum leptocladon is localized on moving sands on the Colorado Plateau. It is not unusual, therefore, for portions of the shrub to be buried in sand, especially the main, woody trunk from which the flowering stems arise. In the Four Corners area three species share portions of overlapping ranges, and herbarium material from there can be difficult to separate. In northwestern New Mexico, var. ramosissimum approaches E. effusum, and in eastern San Juan County the two can be distinguished only by the open, spreading inflorescence branching pattern of var. ramosissimum as compared to the densely branched, compact inflorescence of E. effusum. In the field, the former is confined to sandy areas whereas the latter is typically on heavier, usually clayey soils. In south-central Utah, var. ramosissimum approaches E. nummulare. There, both are found on moving sands, and only the degree of stoutness of the branches can be used to differentiate them when cauline leaves are absent. For the most part, the inflorescence branches of E. nummulare are more rigid and stouter than those of var. ramosissimum, which tend to be flexible and slender. When leaves are present, those of the former are broader and more apically rounded compared to those of the latter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianths pale yellow to yellow
var. leptocladon
1. Perianths white
→ 2
2. Flowering stems and inflorescence branches tomentose, rarely floccose
var. ramosissimum
2. Flowering stems and inflorescence branches glabrous
var. papiliunculi
Source FNA vol. 5, p. 302. FNA vol. 5, p. 249.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. leptocladon var. leptocladon, E. leptocladon var. papiliunculi, E. leptocladon var. ramosissimum
Synonyms E. effusum subsp. leptocladon
Name authority Greene: Bull. Calif. Acad. Sci. 1: 11. (1884) Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(1): 129. (1857)
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