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annual buckwheat, annual wild buckwheat

acorn buckwheat, glandular buckwheat, glandular wild buckwheat

Habit Herbs, 5–20 × 5–10 dm, grayish. Herbs, spreading, annual, (0.5–) 1–2.5 dm, glandular, greenish or reddish green.
Stems

caudex absent;

aerial flowering stems erect, solid, not fistulose, 0.3–0.7 dm, glandular.

Aerial flowering stems

slender, 4–10(–15) dm, floccose to densely tomentose.

Leaves

petiole (rosette) 0.3–1.2 cm, or petiole (cauline) 0.2–0.5 cm, tomentose to floccose;

blade oblanceolate to oblong, 1–7 × 0.3–1.5 cm, densely tomentose abaxially, floccose adaxially not thickened and auriculate-subclasping proximally;

margins entire or slightly revolute.

basal, often sheathing up stem 1–2 cm;

petiole 0.3–2 cm, glandular;

blade broadly elliptic to oval, 0.5–1.5 × 0.5–1.5 cm, pilose-hirsutulous, slightly glandular and greenish on both surfaces, margins entire.

Inflorescences

3–10 × 2–7 cm;

bracts triangular, 1–4 mm.

cymose, diffuse, spreading and usually flat-topped, 5–20 × 5–30 cm;

branches not fistulose, glandular;

bracts 3, scalelike, 0.8–1.2 × 0.5–1 mm.

Peduncles

0.1–0.5 cm, tomentose to floccose.

deflexed or nearly so, straight, slender, 0.2–0.5 cm at proximal nodes, 0.01–0.1 cm distally, sparsely glandular nearly entire length.

Involucres

turbinate to campanulate, 2.5–4 × 2–3 mm, tomentose to floccose abaxially, glabrous adaxially;

teeth 5–6, 0.4–1 mm.

narrowly turbinate, 0.8–1.2(–1.5) × 0.6–1(–1.3) mm, glabrous;

teeth 5, erect, 0.3–0.5 mm.

Flowers

1–2.5 mm;

perianth white or cream to rose;

tepals: those of outer whorl obovate, 1–2 × 0.9–1.5 mm, those of inner whorl narrowly ovate to oblong, 1.5–4 × 1.2–1.8 mm;

stamens usually included, 1–2 mm.

1–1.8 mm;

perianth white to pinkish with dark reddish brown to red midribs, densely pilose;

tepals monomorphic, narrowly lanceolate;

stamens exserted, 1–1.5 mm;

filaments glabrous.

Achenes

1.5–2 mm.

black, 3-gonous, 1–1.3 mm, glabrous.

2n

= 40.

Eriogonum annuum

Eriogonum glandulosum

Phenology Flowering Apr–Nov. Flowering May–Nov.
Habitat Sandy flats, slopes, dunes, and banks, mixed grassland, oak and conifer woodlands Sandy to gravelly, often calcareous slopes, saltbush and creosote bush communities
Elevation (0-)100-1900(-2300) m ((0-)300-6200(-7500) ft) 900-1600 m (3000-5200 ft)
Distribution
from FNA
CO; KS; MT; ND; NE; NM; OK; SD; TX; WY; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Eriogonum annuum is widespread and common to locally abundant or even weedy on the Great Plains of the central United States and extreme north-central Mexico. It was collected in Sherburne County, Minnesota, in 1982, but that population did not persist. The species was recently found as an introduction at Sandy Hook in Monmouth County, New Jersey (Snyder & McArthur s.n., NY), but its fate there remains to be determined. Unfortunately, this weedy species recently has been introduced into northern Arizona as a roadside wild flower. C. L. Perez et al. (1998) have demonstrated that the seed bank can be rich in seeds of this species, but germination rates are low.

The Lakota people traditionally used the annual wild buckwheat as an aid in the treatment of sore mouths in children, seemingly in association with teething (D. J. Rogers 1980). Leaves were used to stain buffalo and deer hides by the Kiowa (P. A. Vestal and R. E. Schultes 1939). Vestal (1952) stated that the species was considered a “life medicine” by the Navajo (Diné) people; it was used also for protection against witches. It is likely that E. annuum was obtained by the Navajo through trade, but it might have been grown locally in historic times where the species recently has been reintroduced.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eriogonum glandulosum is a localized species that varies from rare to common in southeastern Inyo and extreme northeastern San Bernardino counties, California, and in southern Nye, northern Clark, and southwestern Lincoln counties, Nevada. The name E. glandulosum may not be the earliest for the taxon. The type of E. cordatum Torrey & Frémont (1845, as “cordalum”) was collected by Frémont most likely near the California-Nevada border, where E. trichopes, E. contiguum, and E. glandulosum occur. Because the description is not decisive, and because J. Torrey and A. Gray (1870) reported the type as missing, it is impossible to place the name unequivocally. Nonetheless, E. cordatum was reported to be glandular (“glaneous”) with the leaves “pubescent above, hairy underneath.” The glandular condition probably eliminates E. trichopes, while those leaf features probably exclude E. contiguum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5. FNA vol. 5, p. 389.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Micrantha Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Synonyms E. annuum subsp. cymosum, E. annuum subsp. hitchcockii Oxytheca glandulosa, E. carneum, E. glandulosum var. carneum, E. trichopes subsp. glandulosum
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 5: 164. (1835) (Nuttall) Nuttall ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 21. (1856)
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