|
angle stem buckwheat, angle-stem wild buckwheat
|
bastard-sage, bastard-sage buckwheat, Wright's buckwheat, Wright's wild buckwheat
|
| Herbs, erect to spreading, annual, 1–5(–10) dm, tomen-tose to floccose or glabrous, usually grayish. |
Shrubs, subshrubs, or herbs, rarely scapose, (1–)1.5–10 × 1–15(–18) dm or, if matted, 0.1–2.5(–3) × 0.5–3(–5) dm, , lanate to thinly tomentose, or glabrous, grayish to greenish or reddish. |
caudex absent; aerial flowering stems erect, striated, angled, solid, not fistulose, 0.5–1 dm, tomentose to floccose. |
spreading to erect, with or without persistent leaf bases, up to 1/2 or more height of plant; caudex stems absent or spreading, occasionally matted; aerial flowering stems erect to spreading, stout to slender, solid, not fistulose, (0.1–)0.5–4(–6) dm, tomentose, floccose, or glabrous. |
basal and cauline; basal: petiole 0.5–3 cm, mostly floccose, blade oblanceolate to oblong-lanceolate, 1–4(–4.5) × (0.2–)0.5–1(–1.3) cm, tomentose abaxially, floccose or glabrate and grayish or greenish adaxially, margins crenulate; cauline sessile, blade lanceolate to oblong, 0.5–2 × 0.3–0.8 cm, similar to basal blade. |
basal and fasciculate in terminal tufts, or cauline and fasciculate, occasionally 1 per node; petiole 0.02–0.5(–1) cm, tomentose to floccose; blade oblanceolate to broadly elliptic, 0.1–3 × 0.1–1 cm, tomentose to floccose, sometimes subglabrous or glabrous and green adaxially, margins plane, sometimes revolute. |
cymose, open, 5–80 × 10–60 cm; branches striated, angled, sparsely tomentose to glabrate; bracts 3, scalelike, 1–3 × 1–3 mm. |
virgate or cymose with involucres disposed at tips racemosely arranged involucres, rarely capitate, (1–)5–20 × (1–)10–40 cm; branches dichotomous, tomentose, floccose, or glabrous; bracts 3, triangular, scalelike, 0.5–3.5 mm. |
erect, straight, slender, 1–2 cm, sparsely tomentose to glabrous. |
absent. |
turbinate-campanulate to campanulate, 1.5–2.5(–3) × 1.5–2.5(–3), sparsely puberulent; teeth 5, erect, 0.3–0.6 mm. |
1 per node, turbinate to narrowly campanulate, (0.7–)1–4 × 1–2.5 mm, tomentose, floccose, or glabrous; teeth 5, erect, 0.3–1 mm. |
1.5–1.8 mm; perianth white to rose, without a conspicuous rose-purple spot on each outer tepal, minutely glandular-puberulent; tepals dimorphic, those of outer whorl elliptic to obovate, sometimes inflated proximally, those of inner whorl narrowly spatulate; stamens exserted, 2–3 mm; filaments pilose proximally. |
1–4 mm; perianth white to pink or rose, glabrous; tepals connate proximal 1/4, monomorphic, obovate; stamens exserted, 1.5–4 mm; filaments glabrous or sparsely pilose proximally. |
light brown to brown, 3-gonous, 1–1.5 mm, glabrous. |
light brown to brown, (1–)1.5–3 mm, glabrous. |
|
|
|
|
| Flowering year-round. |
|
| Clayey flats and slopes, mixed grassland, saltbush, and chaparral communities, oak and conifer woodlands |
|
| 0-800 m (0-2600 ft) |
|
|
CA
|
AZ; CA; NM; NV; TX; UT; including nw Mexico
|
The name Eriogonum angulosum has been applied to all of the members of its species complex except E. gossypinum. Since the 1950s, the name consistently has been applied to plants with long, exserted stamens and strongly angled stems of the Inner Coast Ranges (Alameda and Contra Costa counties south), the western foothills of the southern Sierra Nevada (Tulare County south), and the Central Valley (San Joaquin County south). The southern edge of the range is the northern foothills of the Transverse Ranges (Ventura and Los Angeles counties). The species can be common and occasionally abundant but rarely is weedy. A mixed collection (with E. gracillimum) from Barstow, San Bernardino County (K. Brandegee s.n., May 1913, UC), and two sheets of the species from San Diego gathered by Susan Stokes apparently in 1895 (B, SD) are discounted as to location. In late fruit, the bractlets at the base of the pedicel inside the involucres of Eriogonum angulosum often elongate and broaden into oblanceolate segments that fill the involucre. As a result, the involucre appears to have several rows of teeth. This feature may be seen also in E. viridescens, but typically the involucres there appear to have only two or three rows of teeth. This feature is seen rarely in E. maculatum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 9 (6 in the flora). Eriogonum wrightii is subdivided into several varieties most of which are distinct, although a few have rather indistinct boundaries. Some of the variation has yet to be fully resolved, especially in the var. nodosum complex where, at least in Mexico, one additional expression remains to be named. A clear distinction between var. subscaposum and E. kennedyi is not possible in southern California (see discussion below). Nearly all of the varieties of E. wrightii are in cultivation, although the most elegant (var. olanchense) has yet to be so honored. Few ethnobotanical uses of bastard-sage are reported in the literature. L. C. Wyman and S. K. Harris (1951) noted that the Kayenta Navajo use it (var. wrightii) as an emetic, while M. L. Zigmond (1981) stated that the Kawaiisu used the pounded seeds (probably of var. subscaposum) in a beverage or as a dry meal. Members of the species are food plants for the rare Rita dotted-blue butterfly (Euphilotes rita), the Pacific dotted-blue (E. enoptes), the veined blue (Plebeius neurona), and the Mormon metalmark (Apodemia mormo mormo). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
1. Plants loosely to compactly matted or herbs | → 2 |
1. Plants shrubs or subshrubs | → 3 |
2. Plants 0.5-2.5(-3) dm; leaf blades 0.5-1(-1.2) cm; involucres 1.5-4 mm; e and s California,wc Nevada | var. subscaposum |
2. Plants 0.1-0.3(-0.6) dm; leaf blades 0.1-0.25 cm; involucres 0.8-1.7(-2) mm; se California | var. olanchense |
3. Flowering stems and branches grayish, lanate to densely tomentose; sw Arizona, se California | var. nodosum |
3. Flowering stems and inflorescence branches whitish, reddish, or greenish, tomentose to floccose; widespread | → 4 |
4. Petiole bases forming distinct ring around stem; leaf blades 0.2-0.6(-1) × 0.1-0.3(-0.4) cm; s California | var. membranaceum |
4. Petiole bases not forming distinct ring around stem; leaf blades 0.5-3 × 0.2-1 cm; sw United States, Mexico | → 5 |
5. Leaf blades 0.5-1.5 × 0.2-0.5(-0.7) cm; involucres 2-2.5 mm; flowers 2.5-3.5 mm; se California to w Texas | var. wrightii |
5. Leaf blades 1.5-3 × 0.5-1 cm; involucres 3-4 mm; flowers 3-4 mm; c and nw California | var. trachygonum |
|
| FNA vol. 5, p. 411. |
FNA vol. 5, p. 289. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. zionis |
|
|
|
E. trachygonum subsp. wrightii |
| Bentham: Trans. Linn. Soc. London 17: 406, plate 18, fig. 1. (1836) |
Torrey ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 14: 15. (1856) |
| |
CA
OR
Flora NW
PNW Herbaria
WildflowerSearch
iNaturalist
USDA Plants Database
