|
angle stem buckwheat, angle-stem wild buckwheat
|
clay-loving buckwheat, clay-loving wild buckwheat
|
| Herbs, erect to spreading, annual, 1–5(–10) dm, tomen-tose to floccose or glabrous, usually grayish. |
Subshrubs, spreading, not scapose, 0.5–1(–1.2) × 0.8–3(–4) dm, floccose or glabrous, grayish. |
caudex absent; aerial flowering stems erect, striated, angled, solid, not fistulose, 0.5–1 dm, tomentose to floccose. |
spreading, without persistent leaf bases, up to 1/3 height of plant; caudex stems absent or compact; aerial flowering stems spreading to erect, slender, solid, not fistulose, 0.05–0.1 dm, thinly floccose or glabrous. |
basal and cauline; basal: petiole 0.5–3 cm, mostly floccose, blade oblanceolate to oblong-lanceolate, 1–4(–4.5) × (0.2–)0.5–1(–1.3) cm, tomentose abaxially, floccose or glabrate and grayish or greenish adaxially, margins crenulate; cauline sessile, blade lanceolate to oblong, 0.5–2 × 0.3–0.8 cm, similar to basal blade. |
cauline, 1 per node; petiole 0.05–0.1 cm, floccose; blade oblanceolate, 0.5–1.2(–1.5) × 0.08–0.2(–0.3) cm, densely white-tomentose abaxially, subglabrous or glabrous and green adaxially, margins tightly revolute. |
cymose, open, 5–80 × 10–60 cm; branches striated, angled, sparsely tomentose to glabrate; bracts 3, scalelike, 1–3 × 1–3 mm. |
cymose, compact, 0.1–2 × 1–3 cm; branches dichotomous, thinly floccose or glabrous; bracts 3, scalelike, triangular, 0.5–1 mm. |
erect, straight, slender, 1–2 cm, sparsely tomentose to glabrous. |
absent or erect, 0.1–0.5 cm, floccose or glabrous. |
turbinate-campanulate to campanulate, 1.5–2.5(–3) × 1.5–2.5(–3), sparsely puberulent; teeth 5, erect, 0.3–0.6 mm. |
1 per node, narrowly turbinate, (2–)2.5–3.5 × 1–1.5 mm, floccose or glabrous; teeth 5, erect, 0.3–0.4 mm. |
1.5–1.8 mm; perianth white to rose, without a conspicuous rose-purple spot on each outer tepal, minutely glandular-puberulent; tepals dimorphic, those of outer whorl elliptic to obovate, sometimes inflated proximally, those of inner whorl narrowly spatulate; stamens exserted, 2–3 mm; filaments pilose proximally. |
(2.5–)3–3.5 mm; perianth cream, glabrous; tepals connate proximal 1/2, essentially monomorphic, oblong; stamens slightly exserted, 2.5–4 mm; filaments sparsely pilose proximally. |
light brown to brown, 3-gonous, 1–1.5 mm, glabrous. |
light brown, 3–3.5 mm, glabrous. |
|
= 40. |
|
|
|
|
| Flowering year-round. |
Flowering May–Jul. |
| Clayey flats and slopes, mixed grassland, saltbush, and chaparral communities, oak and conifer woodlands |
Heavy clay flats and slopes, saltbush communities |
| 0-800 m (0-2600 ft) |
1600-1900 m (5200-6200 ft) |
|
CA
|
CO |
The name Eriogonum angulosum has been applied to all of the members of its species complex except E. gossypinum. Since the 1950s, the name consistently has been applied to plants with long, exserted stamens and strongly angled stems of the Inner Coast Ranges (Alameda and Contra Costa counties south), the western foothills of the southern Sierra Nevada (Tulare County south), and the Central Valley (San Joaquin County south). The southern edge of the range is the northern foothills of the Transverse Ranges (Ventura and Los Angeles counties). The species can be common and occasionally abundant but rarely is weedy. A mixed collection (with E. gracillimum) from Barstow, San Bernardino County (K. Brandegee s.n., May 1913, UC), and two sheets of the species from San Diego gathered by Susan Stokes apparently in 1895 (B, SD) are discounted as to location. In late fruit, the bractlets at the base of the pedicel inside the involucres of Eriogonum angulosum often elongate and broaden into oblanceolate segments that fill the involucre. As a result, the involucre appears to have several rows of teeth. This feature may be seen also in E. viridescens, but typically the involucres there appear to have only two or three rows of teeth. This feature is seen rarely in E. maculatum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eriogonum pelinophilum is a federally listed endangered species with designated critical habitat. It is known only from Mancos Shale hills in Delta and Montrose counties. Much of the former habitat in the Montrose, Colorado, area has been destroyed since the species was listed in 1984. The type locality in Delta County was largely destroyed in 2001 by off-road vehicle activities in the designated critical habitat. A small population is preserved at the Fairview Natural Area east of Montrose. Eriogonum pelinophilum is similar to E. clavellatum although the two are well-separated geographically. It is a smaller plant than E. clavellatum in habit. The flowers of E. clavellatum lack the pronounced, rounded, greenish-red to brownish-red base of the perianth seen in E. pelinophilum, and the tepals are distinctly dimorphic in E. clavellatum whereas they are essentially monomorphic in E. pelinophilum. Ants actively pollinate the flowers, being involved with both self- and cross-pollination. Some 50 additional visitors were found associated with the flowers, but none was confirmed as a pollinator (W. R. Bowlin et al. 1993). The species is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 5, p. 411. |
FNA vol. 5, p. 252. |
| Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
| Bentham: Trans. Linn. Soc. London 17: 406, plate 18, fig. 1. (1836) |
Reveal: Great Basin Naturalist 33: 120. (1973) |
| |
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