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angle stem buckwheat, angle-stem wild buckwheat

mountainmist, Parish's buckwheat, Parish's wild buckwheat

Habit Herbs, erect to spreading, annual, 1–5(–10) dm, tomen-tose to floccose or glabrous, usually grayish. Herbs, spreading, annual, 1–4(–5) dm, glabrous, greenish to reddish brown.
Stems

caudex absent;

aerial flowering stems erect, striated, angled, solid, not fistulose, 0.5–1 dm, tomentose to floccose.

caudex absent;

aerial flowering stems erect, solid, not fistulose, 0.3–1 dm, glabrous, glandular distally.

Leaves

basal and cauline;

basal: petiole 0.5–3 cm, mostly floccose, blade oblanceolate to oblong-lanceolate, 1–4(–4.5) × (0.2–)0.5–1(–1.3) cm, tomentose abaxially, floccose or glabrate and grayish or greenish adaxially, margins crenulate;

cauline sessile, blade lanceolate to oblong, 0.5–2 × 0.3–0.8 cm, similar to basal blade.

basal;

petiole 0.5–2.5 cm, hirsute, slightly winged;

blade spatulate, 2–6 × 0.5–2 cm, hirsute and greenish on both surfaces, margins plane, often ciliate.

Inflorescences

cymose, open, 5–80 × 10–60 cm;

branches striated, angled, sparsely tomentose to glabrate;

bracts 3, scalelike, 1–3 × 1–3 mm.

cymose, usually diffuse, 10–35 × 10–50 cm;

branches not fistulose, glabrous, glandular at nodes;

bracts 3, scalelike, 1–2 × 1–2 mm.

Peduncles

erect, straight, slender, 1–2 cm, sparsely tomentose to glabrous.

spreading, straight, capillary, 0.4–1.5(–2.5) cm, glabrous or sparsely glandular at least proximally.

Involucres

turbinate-campanulate to campanulate, 1.5–2.5(–3) × 1.5–2.5(–3), sparsely puberulent;

teeth 5, erect, 0.3–0.6 mm.

turbinate, 0.5–0.9 × 0.5–0.7 mm, glabrous;

teeth 4, erect or nearly so, 0.3–0.5 mm.

Flowers

1.5–1.8 mm;

perianth white to rose, without a conspicuous rose-purple spot on each outer tepal, minutely glandular-puberulent;

tepals dimorphic, those of outer whorl elliptic to obovate, sometimes inflated proximally, those of inner whorl narrowly spatulate;

stamens exserted, 2–3 mm;

filaments pilose proximally.

0.5–0.9 mm;

perianth pink to red with red midribs, puberulent, becoming white with pink to red midribs;

tepals slightly dimorphic, those of outer whorl ovate and 0.5–0.7 mm wide, those of inner whorl oblong and 0.4–0.5 mm wide;

stamens included, 0.5–0.6 mm;

filaments glabrous.

Achenes

light brown to brown, 3-gonous, 1–1.5 mm, glabrous.

dark brown to blackish, 3-gonous, 1–1.3 mm, glabrous.

2n

= 40.

Eriogonum angulosum

Eriogonum parishii

Phenology Flowering year-round. Flowering Jun–Oct.
Habitat Clayey flats and slopes, mixed grassland, saltbush, and chaparral communities, oak and conifer woodlands Granitic sandy flats and slopes, mixed grassland, chaparral, and sagebrush communities, oak and montane conifer woodlands
Elevation 0-800 m (0-2600 ft) (1000-)1300-3200 m ((3300-)4300-10500 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

The name Eriogonum angulosum has been applied to all of the members of its species complex except E. gossypinum. Since the 1950s, the name consistently has been applied to plants with long, exserted stamens and strongly angled stems of the Inner Coast Ranges (Alameda and Contra Costa counties south), the western foothills of the southern Sierra Nevada (Tulare County south), and the Central Valley (San Joaquin County south). The southern edge of the range is the northern foothills of the Transverse Ranges (Ventura and Los Angeles counties). The species can be common and occasionally abundant but rarely is weedy. A mixed collection (with E. gracillimum) from Barstow, San Bernardino County (K. Brandegee s.n., May 1913, UC), and two sheets of the species from San Diego gathered by Susan Stokes apparently in 1895 (B, SD) are discounted as to location.

In late fruit, the bractlets at the base of the pedicel inside the involucres of Eriogonum angulosum often elongate and broaden into oblanceolate segments that fill the involucre. As a result, the involucre appears to have several rows of teeth. This feature may be seen also in E. viridescens, but typically the involucres there appear to have only two or three rows of teeth. This feature is seen rarely in E. maculatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum parishii occurs from southern Mono County in the White Mountains southward in the Sierra Nevada of Inyo and Tulare counties to scattered desert ranges in San Bernardino (San Bernardino Mountains), Riverside (San Jacinto and Santa Rosa mountains), and San Diego (Laguna Mountains) counties. In Mexico, the species occurs as far south as the Sierra San Pedro Martír in Baja California Norte. A disjunct population from Crown King, Yavapai County, Arizona (Beaty s.n., 6 Sep 1951, CAS) requires confirmation. Possibly it is a recent introduction.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 411. FNA vol. 5, p. 390.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Name authority Bentham: Trans. Linn. Soc. London 17: 406, plate 18, fig. 1. (1836) S. Watson: Proc. Amer. Acad. Arts 17: 379. (1882)
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