Flora of North America species comparison

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Eriogonum angulosum

Eriogonum ovalifolium

angle stem buckwheat, angle-stem wild buckwheat

cushion buckwheat, cushion wild buckwheat, oval-leaf buckwheat, oval-leaf eriogonum
Habit Herbs, erect to spreading, annual, 1–5(–10) dm, tomen-tose to floccose or glabrous, usually grayish. Herbs, forming pulvinate to cespitose maps, usually scapose, 0.2–3 × 0.5–5 dm, floccose to tomentose or lanate.
Stems

caudex absent;

aerial flowering stems erect, striated, angled, solid, not fistulose, 0.5–1 dm, tomentose to floccose.

decumbent to spreading, with persistent leaf bases, up to 1/5 height of plant;

caudex stems matted to spreading;

aerial flowering stems scapelike, spreading to erect, infrequently decumbent to ascending, slender, solid, not fistulose, 0.03–4 dm, floccose to tomentose or lanate or nearly glabrous.
Leaves

basal and cauline;

basal: petiole 0.5–3 cm, mostly floccose, blade oblanceolate to oblong-lanceolate, 1–4(–4.5) × (0.2–)0.5–1(–1.3) cm, tomentose abaxially, floccose or glabrate and grayish or greenish adaxially, margins crenulate;

cauline sessile, blade lanceolate to oblong, 0.5–2 × 0.3–0.8 cm, similar to basal blade.

basal, 1 per node;

petiole not twisted or curled, 0.1–10 cm, mostly tomentose;

blade oblanceolate to elliptic or spatulate to rounded, 0.2–6 × (0.1–)0.2–1.5 cm, lanate to tomentose or floccose, sometimes less so adaxially, margins plane, occasionally brownish.
Inflorescences

cymose, open, 5–80 × 10–60 cm;

branches striated, angled, sparsely tomentose to glabrate;

bracts 3, scalelike, 1–3 × 1–3 mm.

capitate or rarely umbellate, 0.7–5(–7) × 1.5–5 cm;

bracts 3, scalelike, linear to triangular, 0.8–4 mm.
Peduncles

erect, straight, slender, 1–2 cm, sparsely tomentose to glabrous.

absent.
Involucres

turbinate-campanulate to campanulate, 1.5–2.5(–3) × 1.5–2.5(–3), sparsely puberulent;

teeth 5, erect, 0.3–0.6 mm.

1 per node or (2–)3–15 per cluster, turbinate to turbinate-campanulate, (2–)3.5–5(–8) × 2–4 mm, tomentose to floccose;

teeth 5, erect, 0.1–1 mm.
Flowers

1.5–1.8 mm;

perianth white to rose, without a conspicuous rose-purple spot on each outer tepal, minutely glandular-puberulent;

tepals dimorphic, those of outer whorl elliptic to obovate, sometimes inflated proximally, those of inner whorl narrowly spatulate;

stamens exserted, 2–3 mm;

filaments pilose proximally.

(2.5–)3–6(–7) mm;

perianth yellow or white to cream, rose, red, or purple, glabrous;

tepals connate proximally, dimorphic, those of outer whorl usually oval to orbiculate, 2–4 × 2–4 mm, those of inner whorl oblanceolate to elliptic, 3–7 × 0.8–1.5 mm;

stamens mostly included, 1–3 mm;

filaments pilose proximally.
Achenes

light brown to brown, 3-gonous, 1–1.5 mm, glabrous.

light brown to brown, 2–3 mm, glabrous.

Eriogonum angulosum

Eriogonum ovalifolium
Phenology Flowering year-round.
Habitat Clayey flats and slopes, mixed grassland, saltbush, and chaparral communities, oak and conifer woodlands
Elevation 0-800 m (0-2600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

The name Eriogonum angulosum has been applied to all of the members of its species complex except E. gossypinum. Since the 1950s, the name consistently has been applied to plants with long, exserted stamens and strongly angled stems of the Inner Coast Ranges (Alameda and Contra Costa counties south), the western foothills of the southern Sierra Nevada (Tulare County south), and the Central Valley (San Joaquin County south). The southern edge of the range is the northern foothills of the Transverse Ranges (Ventura and Los Angeles counties). The species can be common and occasionally abundant but rarely is weedy. A mixed collection (with E. gracillimum) from Barstow, San Bernardino County (K. Brandegee s.n., May 1913, UC), and two sheets of the species from San Diego gathered by Susan Stokes apparently in 1895 (B, SD) are discounted as to location.

In late fruit, the bractlets at the base of the pedicel inside the involucres of Eriogonum angulosum often elongate and broaden into oblanceolate segments that fill the involucre. As a result, the involucre appears to have several rows of teeth. This feature may be seen also in E. viridescens, but typically the involucres there appear to have only two or three rows of teeth. This feature is seen rarely in E. maculatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 11 (11 in the flora).

Eriogonum ovalifolium is a highly diverse and widespread complex of generally distinct but sometimes intergrading varieties. Several varieties are in cultivation and make worthy additions, especially to the rock garden. The dimorphic nature of the tepals is obvious only in fully mature flowers. In some populations of the more depauperate varieties, such as var. nivale and var. depressum, the tepals may not be as distinctly dimorphic. Nonetheless, the overall aspect of the species is unmistakable.

There are several reports of traditional use of Eriogonum ovalifolium by Native Americans. P. Train et al. (1941) indicated that a decoction of the roots is used in Nevada for colds. R. V. Chamberlin (1911) reported that the Gosiute Indians in southwestern Utah used it in a poultice or wash to treat venereal diseases.

Members of the species are food plants for Bauer’s dotted-blue butterfly (Euphilotes baueri).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Leaf blades 0.2-1.2(-2) cm; scapes usually 0.3-5(-9) cm, rarely longer; involucres usually 2-4.5 (rarely 5-8) mm
→ 2
1. Leaf blades usually 1-6 cm, occasionally shorter; scapes (1-)5-30(-40) cm; involucres (3.5-)4-6.5(-8) mm
→ 8
2. Perianths yellow; c Nevada and ec California
var. caelestinum
2. Perianths white, sometimes rose, purple, or red; widespread
→ 3
3. Leaf margins brownish
→ 4
3. Leaf margins not brownish or, if so, plants of high elevation, central Sierra Nevada
→ 5
4. Leaf blades densely lanate, margins brownish
var. eximium
4. Leaf blades tomentose, margins not brownish
var. williamsiae
5. Involucres 5-8 mm
→ 6
5. Involucres 3-4.5 mm
→ 7
6. Leaf blades white-lanate; San Bernardino Mountains, s California
var. vineum
6. Leaf blades tomentose to floccose; s Sierra Nevada, California
var. monarchense
7. Leaf blades greenish and thinly tomentose at least adaxially, usually elliptic, infrequently oblong to spatulate; scapes often suberect to decumbent, usually thinly floccose; Rocky Mountains of Alberta, Idaho, Montana, and Wyoming, desert ranges of ne Nevada and e Oregon
var. depressum
7. Leaves densely lanate to white-tomentose on both surfaces or only slightly less so adaxially, not at all greenish, usually round; scapes usually erect, lanate or tomentose; Sierra-Cascade cordillera, desert ranges of Great Basin
var. nivale
8. Scapes usually 1-5(-7.5) cm; leaf margins brownish; rare, wc Nevada, ec California
→ 9
8. Scapes usually (4-)7-30(-40) cm, rarely shorter; leaf margins only rarely brownish; widespread
→ 10
9. Leaf blades densely lanate, brownish margins
var. eximium
9. Leaf blades white-tomentose, margins not brownish
var. williamsiae
10. Inflorescences umbellate, branches 1-3 cm; c Idaho and Montana
var. pansum
10. Inflorescences capitate, branches absent; widespread
→ 11
11. Flowers 5-7 mm; involucres 5-7 mm; s California
var. vineum
11. Flowers 4-5 mm; involucres 4-6.5 mm; widespread
→ 12
12. Perianths yellow
var. ovalifolium
12. Perianths white to rose or purple.
→ 13
13. Leaf blades spatulate, oblong, or obovate to oval; scapes (4-)5-20 cm; widespread
var. purpureum
13. Leaf blades oblanceolate to narrowly elliptic; scapes (15-)20-30(-40) cm; Montana and Wyoming
var. ochroleucum
Source FNA vol. 5, p. 411. FNA vol. 5, p. 319.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. ovalifolium var. caelestinum, E. ovalifolium var. depressum, E. ovalifolium var. eximium, E. ovalifolium var. monarchense, E. ovalifolium var. nivale, E. ovalifolium var. ochroleucum, E. ovalifolium var. ovalifolium, E. ovalifolium var. pansum, E. ovalifolium var. purpureum, E. ovalifolium var. vineum, E. ovalifolium var. williamsiae
Synonyms Eucycla ovalifolia
Name authority Bentham: Trans. Linn. Soc. London 17: 406, plate 18, fig. 1. (1836) Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 50, plate 8, fig. 1. (1834)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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