Flora of North America species comparison

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Eriogonum angulosum

Eriogonum nudum

angle stem buckwheat, angle-stem wild buckwheat

bare-stem buckwheat, barestem wild buckwheat, naked buckwheat, naked wild buckwheat, nude buckwheat
Habit Herbs, erect to spreading, annual, 1–5(–10) dm, tomen-tose to floccose or glabrous, usually grayish. Herbs, mostly erect, infrequently scapose, (0.5–)1–15(–20) × 0.5–3 dm, glabrous or floccose to tomentose, usually greenish, occasionally grayish.
Stems

caudex absent;

aerial flowering stems erect, striated, angled, solid, not fistulose, 0.5–1 dm, tomentose to floccose.

spreading to erect, with or without persistent leaf bases, up to 1/4 height of plant;

caudex stems absent;

aerial flowering stems erect to spreading, slender to stout, solid or hollow, occasionally fistulose, 0.3–4(–10) dm, glabrous or lanate to tomentose or floccose.
Leaves

basal and cauline;

basal: petiole 0.5–3 cm, mostly floccose, blade oblanceolate to oblong-lanceolate, 1–4(–4.5) × (0.2–)0.5–1(–1.3) cm, tomentose abaxially, floccose or glabrate and grayish or greenish adaxially, margins crenulate;

cauline sessile, blade lanceolate to oblong, 0.5–2 × 0.3–0.8 cm, similar to basal blade.

basal or sheathing up stem 0.5–4 dm;

petiole 1–10 cm, glabrous or tomentose;

blade oblanceolate to elliptic or ovate, 1–6 × (0.3–)1–4 cm, densely white-lanate or tomentose abaxially, tomentose to floccose or subglabrous to glabrous adaxially, margins plane or undulate-crisped.
Inflorescences

cymose, open, 5–80 × 10–60 cm;

branches striated, angled, sparsely tomentose to glabrate;

bracts 3, scalelike, 1–3 × 1–3 mm.

cymose, rarely umbellate or capitate, 2–100(–150) × 2–40(–80) cm;

branches usually dichotomous, glabrous or tomentose to floccose or sparsely pubescent;

bracts 3, scalelike, 0.5–3(–5) mm.
Peduncles

erect, straight, slender, 1–2 cm, sparsely tomentose to glabrous.

absent.
Involucres

turbinate-campanulate to campanulate, 1.5–2.5(–3) × 1.5–2.5(–3), sparsely puberulent;

teeth 5, erect, 0.3–0.6 mm.

1 per node or 2–10 per cluster, turbinate to turbinate-campanulate, (2.5–)3–5(–7) × (1.5–)2–4 mm, glabrous, tomentose, or sparsely pubescent;

teeth 5–8, 0.2–0.6 mm.
Flowers

1.5–1.8 mm;

perianth white to rose, without a conspicuous rose-purple spot on each outer tepal, minutely glandular-puberulent;

tepals dimorphic, those of outer whorl elliptic to obovate, sometimes inflated proximally, those of inner whorl narrowly spatulate;

stamens exserted, 2–3 mm;

filaments pilose proximally.

(1.5–)2–4 mm;

perianth white or yellow, sometimes pink or rose, glabrous or pubescent;

tepals connate proximal 1/4, monomorphic, oblong to obovate;

stamens exserted, 2–5 mm;

filaments pilose proximally.
Achenes

light brown to brown, 3-gonous, 1–1.5 mm, glabrous.

light brown to brown, 1.5–3.5 mm, glabrous.

Eriogonum angulosum

Eriogonum nudum
Phenology Flowering year-round.
Habitat Clayey flats and slopes, mixed grassland, saltbush, and chaparral communities, oak and conifer woodlands
Elevation 0-800 m (0-2600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR; WA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

The name Eriogonum angulosum has been applied to all of the members of its species complex except E. gossypinum. Since the 1950s, the name consistently has been applied to plants with long, exserted stamens and strongly angled stems of the Inner Coast Ranges (Alameda and Contra Costa counties south), the western foothills of the southern Sierra Nevada (Tulare County south), and the Central Valley (San Joaquin County south). The southern edge of the range is the northern foothills of the Transverse Ranges (Ventura and Los Angeles counties). The species can be common and occasionally abundant but rarely is weedy. A mixed collection (with E. gracillimum) from Barstow, San Bernardino County (K. Brandegee s.n., May 1913, UC), and two sheets of the species from San Diego gathered by Susan Stokes apparently in 1895 (B, SD) are discounted as to location.

In late fruit, the bractlets at the base of the pedicel inside the involucres of Eriogonum angulosum often elongate and broaden into oblanceolate segments that fill the involucre. As a result, the involucre appears to have several rows of teeth. This feature may be seen also in E. viridescens, but typically the involucres there appear to have only two or three rows of teeth. This feature is seen rarely in E. maculatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 13 (13 in the flora).

Several local groups of Native Americans in California used members of this species in a variety of ways. S. A. Barrett and E. W. Gifford (1933) and S. M. Schenck and E. W. Gifford (1952) reported the consumption of raw young stems that are rather moist and tasty, although there is a sour aftertaste. The Kawaiisu used the hollow stems (probably var. westonii, rather than var. pauciflorum) as drinking tubes and as pipes (M. L. Zigmond 1981). Zigmond reported also that the roots of var. pauciflorum are used as an infusion for coughs.

Members of Eriogonum nudum are food plants for the Bauer’s dotted-blue butterfly (Euphilotes baueri), the Pacific dotted-blue (E. enoptes), the gorgon copper (Gaeides gorgon), and the Mormon metalmark (Apodemia mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Involucres and inflorescence branches tomentose to floccose; leaves usually basal
→ 2
1. Involucres and inflorescence branches glabrous or, if pubescent, leaves sheathing up stems
→ 3
2. Perianths white; flowering stems lanate to tomentose; leaf blades 2-3.5 cm; s Sierra Nevada
var. regirivum
2. Perianths yellow or white to rose; flowering stems tomentose to floccose; leaf blades 2-4 cm; s Oregon, n California, wc Nevada
var. oblongifolium
3. Leaves sheathing up stems, margins often strongly undulate-crisped
→ 4
3. Leaves basal, margins plane or slightly undulate-crisped
→ 8
4. Flowering stems tomentose
→ 5
4. Flowering stems glabrous
→ 6
5. Perianths pubescent; flowers 1.5-2 mm; involucres 3-4 mm; Sierra Nevada, California
var. regirivum
5. Perianths glabrous; flowers 3-4 mm; involucres 4-6 mm; Coast Ranges, California
var. decurrens
6. Leaf blades densely lanate abaxially, tomentose adaxially; involucres 5-10 per cluster; Sierra Nevada, California
var. murinum
6. Leaf blades tomentose abaxially, less so to floccose, glabrous, or nearly so adaxially; involucres 1 per node or 2-5 per cluster; widespread, Coast Ranges, California
→ 7
7. Flowering stems occasionally fistulose; involucres (2-)3-5 per cluster; perianths white to pink, rarely yellowish
var. auriculatum
7. Flowering stems strongly fistulose; involucres 1 per node; perianths pale yellowish white to yellow or white
var. indictum
8. Involucres 1(-2) per cluster
var. westonii
8. Involucres 2-10 per cluster
→ 12
9. Flowering stems fistulose; perianths yellow, infrequently white; c California
var. westonii
9. Flowering stems not fistulose or, if so, plants of sw California; perianths white, rarely yellow
→ 10
10. Perianths pubescent or, infrequently, glabrous; s California
var. pauciflorum
10. Flowers glabrous; sw Oregon, n California, Nevada
→ 11
11. Leaf blades 1-5 cm; sw Oregon, n California (including w slope of Sierra Nevada), Washington
var. nudum
11. Leaf blades 1-2 cm; Sierra Nevada of California and adjacent desert ranges of wc Nevada
var. deductum
12. Perianths pubescent, often yellow
→ 13
12. Perianths usually glabrous abaxially, white, rarely yellow
→ 14
13. Flowering stems not fistulose; leaf blades floccose or glabrous adaxially, margins plane; plants of nonaridregions
var. pubiflorum
13. Flowering stems slightly to distinctly fistulose; leaf blades tomentose to floccose adaxially, margins undulate-crisped; plants of arid regions
var. westonii
14. Inflorescences capitate or nearly so; alpine, Sierra Nevada
E. nudumvar. scapigerum
14. Inflorescences cymose or, if capitate, not alpine.
→ 15
15. Involucres 1(-2) per cluster, 5- 7 mm; mountains of s California
var. pauciflorum
15. Involucres 2-10 per cluster, 3-5 mm; mountains and foothills of c and n California or coastal bluffs. [16. Shifted to left margin—Ed.]
→ 16
16. Inflorescences cymose and branched 2 or more times; involucres 2-5 per cluster; mountains and foothills of c and n California, Oregon, and Washington.
var. nudum
16. Inflorescences capitate or cymose and branched 1-2 times; involucres 5-10 per cluster; coastal bluffs, sw Oregon, ne California
var. paralinum
Source FNA vol. 5, p. 411. FNA vol. 5, p. 309.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. nudum var. auriculatum, E. nudum var. decurrens, E. nudum var. deductum, E. nudum var. indictum, E. nudum var. murinum, E. nudum var. nudum, E. nudum var. oblongifolium, E. nudum var. paralinum, E. nudum var. pauciflorum, E. nudum var. pubiflorum, E. nudum var. regirivum, E. nudum var. westonii, E. nudumvar. scapigerum
Synonyms E. latifolium subsp. nudum
Name authority Bentham: Trans. Linn. Soc. London 17: 406, plate 18, fig. 1. (1836) Douglas ex Bentham: Trans. Linn. Soc. London 17: 413. (1836)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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