angle stem buckwheat, angle-stem wild buckwheat
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slender buckwheat, slender wild buckwheat
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Herbs, erect to spreading, annual, 1–5(–10) dm, tomen-tose to floccose or glabrous, usually grayish. |
Subshrubs or shrubs, erect to spreading, not scapose, 0.2–1.5 × (0.6–)1–13(–16) dm, white- to tannish-tomentose, floccose, or glabrous. |
caudex absent; aerial flowering stems erect, striated, angled, solid, not fistulose, 0.5–1 dm, tomentose to floccose. |
spreading to erect, typically without persistent leaf bases, up to 1/2 height of plant; caudex stems absent or spreading; aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.05–1.5 dm, lanate, tomentose, floccose, subglabrous, or glabrous. |
basal and cauline; basal: petiole 0.5–3 cm, mostly floccose, blade oblanceolate to oblong-lanceolate, 1–4(–4.5) × (0.2–)0.5–1(–1.3) cm, tomentose abaxially, floccose or glabrate and grayish or greenish adaxially, margins crenulate; cauline sessile, blade lanceolate to oblong, 0.5–2 × 0.3–0.8 cm, similar to basal blade. |
cauline, 1 per node or fasciculate; petiole 0.1–0.5 cm, tomentose to floccose or glabrous; blade usually elliptic, sometimes linear to obovate, 0.3–3.5 × (0.07–)0.1–1.2 cm, tomentose abaxially, less so or glabrous adaxially, margins occasionally revolute. |
cymose, open, 5–80 × 10–60 cm; branches striated, angled, sparsely tomentose to glabrate; bracts 3, scalelike, 1–3 × 1–3 mm. |
cymose, compact, often flat-topped, 0.5–6(–12) × 1–10(–13) cm; branches dichotomous, whitish-lanate to brownish- or reddish-tomentose to floccose or glabrate, infrequently green or gray and subglabrous or glabrous; bracts 3, scalelike, linear to triangular, 1–5 mm. |
erect, straight, slender, 1–2 cm, sparsely tomentose to glabrous. |
absent or mostly erect, slender, 0.3–1.5 cm, tomentose to floccose. |
turbinate-campanulate to campanulate, 1.5–2.5(–3) × 1.5–2.5(–3), sparsely puberulent; teeth 5, erect, 0.3–0.6 mm. |
1 per node, turbinate, (1.5–)2–3.5(–4) × 1.3–2.5(–3) mm, tomentose, floccose, subglabrous, or glabrous; teeth 5, erect, (0.3–)0.5–1(–1.7) mm. |
1.5–1.8 mm; perianth white to rose, without a conspicuous rose-purple spot on each outer tepal, minutely glandular-puberulent; tepals dimorphic, those of outer whorl elliptic to obovate, sometimes inflated proximally, those of inner whorl narrowly spatulate; stamens exserted, 2–3 mm; filaments pilose proximally. |
1.5–3(–4) mm; perianth yellow or white to pink, orange, rose, red, or occasionally cream, glabrous; tepals connate proximal 1/5–2/5, essentially monomorphic, oblong to obovate; stamens usually exserted, 2.5–4 mm; filaments sparsely to densely puberulent proximally. |
light brown to brown, 3-gonous, 1–1.5 mm, glabrous. |
brown, 1.5–3 mm, glabrous. |
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Flowering year-round. |
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Clayey flats and slopes, mixed grassland, saltbush, and chaparral communities, oak and conifer woodlands |
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0-800 m (0-2600 ft) |
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CA
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AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY
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The name Eriogonum angulosum has been applied to all of the members of its species complex except E. gossypinum. Since the 1950s, the name consistently has been applied to plants with long, exserted stamens and strongly angled stems of the Inner Coast Ranges (Alameda and Contra Costa counties south), the western foothills of the southern Sierra Nevada (Tulare County south), and the Central Valley (San Joaquin County south). The southern edge of the range is the northern foothills of the Transverse Ranges (Ventura and Los Angeles counties). The species can be common and occasionally abundant but rarely is weedy. A mixed collection (with E. gracillimum) from Barstow, San Bernardino County (K. Brandegee s.n., May 1913, UC), and two sheets of the species from San Diego gathered by Susan Stokes apparently in 1895 (B, SD) are discounted as to location. In late fruit, the bractlets at the base of the pedicel inside the involucres of Eriogonum angulosum often elongate and broaden into oblanceolate segments that fill the involucre. As a result, the involucre appears to have several rows of teeth. This feature may be seen also in E. viridescens, but typically the involucres there appear to have only two or three rows of teeth. This feature is seen rarely in E. maculatum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 13 (13 in the flora). Eriogonum microthecum is used as browse by deer and to a lesser degree by cattle and sheep. Some forms are now in cultivation. The species is reportedly used by the Piute of Nevada in the treatment of tuberculosis, lameness, rheumatism, and bladder trouble (P. Train et al. 1941). S. A. Weber and P. D. Seaman (1985) stated that A. F. Whiting found the plants being used as a tea by the Havasupai in northern Arizona. Members of E. microthecum are food plants for subspecies of the rare pallid blue butterfly (Euphilotes pallescens). Also found on this species is the cythera metalmark (Apodemia mormo cythera). Some authors have referred E. effusum to this species, even though the ranges of the two species do not overlap and intermediates are unknown. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| → 2 |
1. Perianths various shades of white, cream, orange, pink, or red, not yellow | → 4 |
2. Flowering stems and inflorescence branches usually glabrous; e Oregon and wc Idaho | var. microthecum |
2. Flowering stems and inflorescence branches tomentose to floccose; se Oregon and sw Idaho s to e California and w Nevada | → 3 |
3. Leaf blades (0.2-)0.3-0.6(-0.8) cm wide; flowers (1.5-)2-2.5(-3) mm; involucres 2-2.5 mm; achenes 1.5-2 mm; se Oregon and sw Idaho s to e California and w Nevada | var. ambiguum |
3. Leaf blades 0.5-1.2 cm wide; flowers 2.5-3 mm; involucres 2.5-4 mm; achenes 2.5-3 mm; ne California, nw Nevada | var. schoolcraftii |
4. Tomentum whitish (see also var. alpinum of the Sierra Nevada, California); flowering stems and inflorescence branches infrequently glabrous | → 5 |
4. Tomentum brownish or reddish (may be white in var. alpinum), or flowering stems and inflorescence branches essentially glabrous | → 6 |
5. Leaf margins not revolute; flowering inflorescence branches floccose or glabrous; northern phase of species | var. laxiflorum |
5. Leaf margins revolute or nearly so; flowering inflorescence branches lanate to tomentose, or if subglabrous or glabrous, then southern phase of species | var. simpsonii |
| → 7 |
6. Plants subshrubs, 0.2-1.5(-2) dm | → 8 |
7. Flowering stems and inflorescence branches tomentose when young, becoming floccose at maturity; flowers 1.5-2(-2.5) mm; achenes 1.8-2 mm; Death Valley region, California | var. panamintense |
7. Flowering stems and inflorescence branches lanate to tomentose at maturity; flowers 2-2.5(-3) mm; achenes 2.5-3 mm; Transverse Ranges, California | E. microthecumvar. corymbosoides |
8. Leaf blades elliptic or ovate, margins not revolute; flowers (1.5-)2-3.5(-4) mm | → 9 |
8. Leaf blades linear or linear-oblanceolate to narrowly elliptic, margins often revolute; flowers 1.5-2.5 mm | → 10 |
9. Leaf blades 0.5-1 × (0.2-)0.3-0.5(-0.6) cm; involucres (2-)2.5-3 mm; flowers (2.5-)3-3.5(-4) mm; San Gabriel Mountains, California | var. johnstonii |
9. Leaf blades 0.3-0.7(-0.8) × 0.1-0.4 cm; involucres 2.5-3.5 mm; flowers (1.5-)2-3.5 mm; desert ranges of se California, c Nevada, and Utah1f. Eriogonum microthecum var. lapidicola [8. Shifted to left margin—Ed.] | → 8 |
10. Involucres 3-4 mm; perianths cream; San Bernardino Mountains, California | var. lacus-ursi |
10. Involucres (1.5-)2-3 mm; perianths white, pink, red, or rose; Sierra Nevada of California, or e Nevada and w Utah | → 11 |
11. Flowering stems glabrous; leaf blades sparsely floccose or glabrous adaxially; desert ranges, se Nevada | var. arceuthinum |
11. Flowering stems white- to brownish-floccose to subglabrous, or reddish-tomentose to floccose; leaf blades floccose to subglabrous adaxially; California or wc Utah | → 12 |
12. Tomentum white to brownish; Sierra Nevada, California | var. alpinum |
12. Tomentum reddish; desert ranges, wc Utah | var. phoeniceum |
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FNA vol. 5, p. 411. |
FNA vol. 5, p. 242. |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Ganysma |
Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis |
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E. microthecum var. alpinum, E. microthecum var. ambiguum, E. microthecum var. arceuthinum, E. microthecum var. johnstonii, E. microthecum var. lacus-ursi, E. microthecum var. laxiflorum, E. microthecum var. microthecum, E. microthecum var. panamintense, E. microthecum var. phoeniceum, E. microthecum var. schoolcraftii, E. microthecum var. simpsonii, E. microthecumvar. corymbosoides |
Bentham: Trans. Linn. Soc. London 17: 406, plate 18, fig. 1. (1836) |
Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 15. (1848) |
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