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wing buckwheat, wing wild buckwheat

Shockley's buckwheat, Shockley's wild buckwheat

Habit Herbs, monocarpic, 5–20(–25) dm, strigose or glabrous; taproot often chambered. Herbs, matted, scapose, 0.3–0.5(–0.7) × (0.5–)1–4(–20) dm, floccose to tomentose, greenish or grayish.
Stems

caudex absent;

aerial flowering stems usually 1, not fistulose, 2–13 dm, strigose or glabrous.

matted, sometimes only seemingly so, with persistent leaf bases, up to 1/5 height of plant;

caudex stems matted;

aerial flowering stems absent or scapelike, erect or nearly so, slender, solid, not fistulose, (0.05–)0.1–0.3 dm, floccose to tomentose.

Leaves

basal and sometimes cauline;

basal petiolate, petiole 2–6 cm, stigose to woolly or glabrous, blade linear-lanceolate or lanceolate to oblanceolate to spatulate, (3–)5–20 × 0.3–2 cm, strigose, becoming glabrous and green on both surfaces except for margins and midvein;

cauline sessile, blade linear-oblanceolate to lanceolate, 1–9 × 0.3–0.8(–1) cm, similar to basal blade.

basal, fasciculate in terminal tufts;

petiole 0.2–0.5 cm, tomentose to floccose;

blade oblanceolate to elliptic or spatulate, (0.2–)0.3–0.8(–1.2) × 0.2–0.4(–0.6) cm, tomentose to floccose, margins plane or slightly thickened.

Inflorescences

2–10 dm;

branches strigose or glabrous;

bracts semileaflike proximally, linear to linear-lanceolate, 2–9 × 1–3 mm, scalelike distally, triangular, 0.8–5 × 0.5–2 mm.

capitate, 0.8–2 cm;

branches absent;

bracts 3–5, scalelike, linear to linear-lanceolate, 1.5–4 × 0.6–1 mm.

Peduncles

erect, straight or curving upward, 0.5–3.5 cm, strigose or glabrous.

absent.

Involucres

turbinate to campanulate, 2–4(–4.5) × 2–4(–4.5) mm, strigose or glabrous;

teeth 1–1.8 mm.

2–4(–6) per cluster, campanulate, (2–)2.5–5(–6) × 3–6(–7) mm, rigid, tomentose;

teeth 5–10, erect to spreading, (0.5–)1–3 mm.

Flowers

1.5–2.5 mm in anthesis, 3–6 mm in fruit;

perianth yellow to yellowish green, rarely maroon in anthesis, often reddish or maroon in fruit, glabrous;

tepals lanceolate;

stamens 1.5–3 mm;

filaments glabrous.

2.5–4 mm;

perianth white to rose or yellow, densely pilose;

tepals connate proximally, monomorphic, oblong to obovate;

stamens exserted, 2.5–5 mm;

filaments subglabrous or sparsely pilose proximally.

Achenes

yellowish green to reddish brown, 5–9 mm, glabrous, 3-winged entire length, beakless.

light brown to brown, 2.5–3 mm, tomentose.

Eriogonum alatum

Eriogonum shockleyi

Phenology Flowering May–Aug.
Habitat Gravelly or clayey (rarely sandy) flats, washes, and slopes, saltbush, blackbrush, and sagebrush communities, pinyon-juniper woodlands
Elevation (800-)1200-2600 m ((2600-)3900-8500 ft)
Distribution
from FNA
AZ; CO; KS; NE; NM; OK; TX; UT; WY; including Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The mature reddish roots of Eriogonum alatum can be distinctively chambered. The Navajo (Diné) people consider the species to be a “life medicine” (L. C. Wyman and S. K. Harris 1951), using a mixture of shredded roots and water primarily to treat internal ailments. The species is used also as a ceremonial medicine (P. A. Vestal 1952). The Zuni use it as an emetic for stomachaches (S. Camazine and R. A. Bye 1980).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eriogonum shockleyi is widely distributed in northern Arizona, east-central California, western Colorado, southern Idaho, northwestern New Mexico, Nevada, and Utah. On the Colorado Plateau, it has oblanceolate to spatulate leaf blades usually 0.3–1.2 × 0.3–0.6 cm, scapes 1–3 cm, involucres with long (2–3.5 mm) often spreading teeth, and flowers 3–4 mm. These plants have been distinguished as var. longilobum. The typical Great Basin expression has elliptic leaf blades 0.3–0.6 × 0.3–0.5 cm, scapes 0.5–2 cm, involucres with short (0.5–2 mm) erect teeth, and flowers 2.5–4 mm. Low, compact, hummock-like plants in southwestern Idaho with elliptic leaf blades 0.1–0.3(–0.35) × 0.1–0.15 cm, flowering stems absent or up to 0.5 cm, and involucres 2.5–3 mm with teeth 0.8–1 mm have been termed var. packardiae. These morphologic differences have been shown to be genetically insignificant, however (J. F. Smith and T. A. Bateman 2002). Plants on moving sand dunes at the southeast end of Baking Power Flat in Lincoln County, Nevada, can be up to 2 m across. Further studies may indicate that the various geographic expressions deserve taxonomic recognition.

The Great Basin expression is the food plant of the Bernardino dotted-blue butterfly (Euphilotes bernardino). Members of the species are occasionally found in cultivation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowering stems and inflorescence branches usually strigose; peduncles strigose or nearly so; involucres strigose, rarely glabrous; plants 5-13(-17) dm; n Arizona, Colorado, w Kansas, sw Nebraska, New Mexico, w Texas, e Utah, se Wyoming
var. alatum
1. Flowering stems and inflorescence branches glabrous or nearly so; peduncles glabrous, occasionally slightly strigose; involucres glabrous; plants 10-20(-25) dm; ne New Mexico, w Oklahoma, n Texas
var. glabriusculum
Source FNA vol. 5, p. 379. FNA vol. 5, p. 286.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Pterogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. alatum var. alatum, E. alatum var. glabriusculum
Synonyms Pterogonum alatum E. pulvinatum, E. shockleyi subsp. candidum, E. shockleyi subsp. longilobum, E. shockleyi var. longilobum, E. shockleyi var. packardiae, E. villiflorum var. candidum
Name authority Torrey: in L. Sitgreaves, Rep. Exped. Zuni Colorado Rivers, 168, plate 8. (1853) S. Watson: Proc. Amer. Acad. Arts 18: 194. (1883)
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