Flora of North America species comparison

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Eriogonum alatum

Eriogonum heermannii

wing buckwheat, wing wild buckwheat

Heerman buckwheat, Heermann's buckwheat, Heermann's wild buckwheat
Habit Herbs, monocarpic, 5–20(–25) dm, strigose or glabrous; taproot often chambered. Shrubs and subshrubs, spreading to rounded and occasionally erect, not scapose, (0.5–)1–20 × 2–25 dm, glabrous or occasionally floccose, sometimes scabrellous, greenish, infrequently grayish.
Stems

caudex absent;

aerial flowering stems usually 1, not fistulose, 2–13 dm, strigose or glabrous.

spreading or erect, without persistent leaf bases, up to 1/2 height of plant;

caudex stems absent;

aerial flowering stems erect or nearly so, slender to stout, solid, not fistulose, 0.02–0.5 dm, thinly tomentose or glabrous.
Leaves

basal and sometimes cauline;

basal petiolate, petiole 2–6 cm, stigose to woolly or glabrous, blade linear-lanceolate or lanceolate to oblanceolate to spatulate, (3–)5–20 × 0.3–2 cm, strigose, becoming glabrous and green on both surfaces except for margins and midvein;

cauline sessile, blade linear-oblanceolate to lanceolate, 1–9 × 0.3–0.8(–1) cm, similar to basal blade.

cauline, 1 per node, quickly deciduous;

petiole 0.1–1.5 cm, floccose or glabrous;

blade linear, oblanceolate or spatulate or elliptic, or oblong, (0.4–)1–2(–4) × 0.1–0.8 cm, tomentose to floccose or glabrous abaxially, floccose to thinly floccose or glabrous adaxially, margins plane.
Inflorescences

2–10 dm;

branches strigose or glabrous;

bracts semileaflike proximally, linear to linear-lanceolate, 2–9 × 1–3 mm, scalelike distally, triangular, 0.8–5 × 0.5–2 mm.

cymose or racemose, 1–25(–30) × 1–30(–35) cm;

branches dichotomous, sometimes with secondaries suppressed, smooth or angled to ridged and grooved, glabrous or occasionally floccose or scabrous;

bracts 3, scalelike, 0.3–2 mm.
Peduncles

erect, straight or curving upward, 0.5–3.5 cm, strigose or glabrous.

absent.
Involucres

turbinate to campanulate, 2–4(–4.5) × 2–4(–4.5) mm, strigose or glabrous;

teeth 1–1.8 mm.

1 per node, narrowly turbinate or campanulate, 0.7–3 × 0.7–4 mm, glabrous, infrequently floccose;

teeth 5, erect, 0.3–0.7 mm.
Flowers

1.5–2.5 mm in anthesis, 3–6 mm in fruit;

perianth yellow to yellowish green, rarely maroon in anthesis, often reddish or maroon in fruit, glabrous;

tepals lanceolate;

stamens 1.5–3 mm;

filaments glabrous.

(1.5–)2–4 mm;

perianth white, yellowish white, pink, or reddish, glabrous;

tepals connate proximal 1/4, dimorphic, those of outer whorl obovate to orbiculate, those of inner whorl narrowly lanceolate to oblong;

stamens exserted, 2–5 mm;

filaments pilose proximally.
Achenes

yellowish green to reddish brown, 5–9 mm, glabrous, 3-winged entire length, beakless.

light brown to brown, 2–5 mm, glabrous.

Eriogonum alatum

Eriogonum heermannii
Distribution
from FNA
AZ; CO; KS; NE; NM; OK; TX; UT; WY; including Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The mature reddish roots of Eriogonum alatum can be distinctively chambered. The Navajo (Diné) people consider the species to be a “life medicine” (L. C. Wyman and S. K. Harris 1951), using a mixture of shredded roots and water primarily to treat internal ailments. The species is used also as a ceremonial medicine (P. A. Vestal 1952). The Zuni use it as an emetic for stomachaches (S. Camazine and R. A. Bye 1980).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 8 (8 in the flora).

As Eriogonum heermannii is here circumscribed, the number of varieties is dramatically decreased from past presentations, with E. apachense reduced to synonymy under a now greatly expanded var. argense. Also included in that variety is the more stoutly branched var. subracemosum. The southern var. heermannii of basically desert ranges and the more northern var. occidentale of the Coast Ranges in California are maintained, but their separation is more traditional than certain. The fragile and bulky nature of many dried, often poorly prepared specimens, and the tendency for leaves to fall away have made varietal identification within E. heermannii difficult.

Eriogonum heermannii varieties are food plants for Ellis’s dotted-blue butterfly (Euphilotes ellisi) and the Mormon metalmark (Apodemia mormo mormo).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Flowering stems and inflorescence branches usually strigose; peduncles strigose or nearly so; involucres strigose, rarely glabrous; plants 5-13(-17) dm; n Arizona, Colorado, w Kansas, sw Nebraska, New Mexico, w Texas, e Utah, se Wyoming
var. alatum
1. Flowering stems and inflorescence branches glabrous or nearly so; peduncles glabrous, occasionally slightly strigose; involucres glabrous; plants 10-20(-25) dm; ne New Mexico, w Oklahoma, n Texas
var. glabriusculum
1. Flowering stems and inflorescence branches round or angled, scabrellous or papillate-scabrous; plants often densely branched
→ 2
1. Flowering stems and inflorescence branches round, not angled, usually smooth, glabrous or thinly tomentose; plants sparsely branched
→ 3
2. Flowering stems and inflorescence branches round, usually distinctly scabrellous or infrequently papillate-scabrous; Arizona, se California, Nevada
var. argense
2. Flowering stems and inflorescence branches sharply ridged and deeply grooved, minutely scabrellous; nw Arizona, se California, s Nevada, sw Utah
var. sulcatum
3. Involucres not racemosely arranged on inflorescence branches, or only last 2-3 so disposed; inflorescences diffusely branched, glabrous; Great Basin or northeast edge of Mojave Desert
→ 4
3. Involucres racemosely arranged along inflorescence branches or at least at tips of branches; inflorescences openly branched, glabrous or floccose to thinly tomentose; Mojave and Sonoran deserts
→ 5
4. Inflorescence branches mostly slender, smooth, not ridged, not spinose; inflorescences 3-15(-23) × 5-20 cm; subshrubs or shrubs, 3-7 × 5-12(-15) dm; Great Basin, ec California and Nevada
var. humilius
4. Inflorescence branches stoutish, faintly ridged and grooved, spinose; inflorescences 3-7(-10) × 3-10(-12) cm; shrubs, 1-3 × 1.5-5(-8) dm; Mojave Desert, sw Utah and nw Arizona
var. subspinosum
5. Inflorescence branches thinly tomentose to floccose; s California, s Nevada, nw Arizona
var. floccosum
5. Inflorescence branches glabrous; s Nevada, California
→ 6
6. Involucres racemosely arranged; inflorescence branches whiplike; s Nevada
var. clokeyi
6. Involucres racemosely arranged only distally; inflorescence branches not whiplike; California
→ 7
7. Leaf blades 0.5-1.5 cm, glabrous abaxially; inflorescence branches stout; sc California
var. heermannii
7. Leaf blades 1.5-3(-4) cm, mostly tomentose to floccose abaxially (at least in early anthesis); inflorescence branches slender; sw California
var. occidentale
Source FNA vol. 5, p. 379. FNA vol. 5, p. 304.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Pterogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. alatum var. alatum, E. alatum var. glabriusculum
E. heermannii var. argense, E. heermannii var. clokeyi, E. heermannii var. floccosum, E. heermannii var. heermannii, E. heermannii var. humilius, E. heermannii var. occidentale, E. heermannii var. subspinosum, E. heermannii var. sulcatum
Synonyms Pterogonum alatum E. geniculatum
Name authority Torrey: in L. Sitgreaves, Rep. Exped. Zuni Colorado Rivers, 168, plate 8. (1853) Durand & Hilgard: in War Department [U.S.], Pacif. Railr. Rep. 5(3): 14. (1857)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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