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wing buckwheat, wing wild buckwheat

California buckwheat, California wild buckwheat, eastern Mojave buckwheat, eastern Mojave wild buckwheat

Habit Herbs, monocarpic, 5–20(–25) dm, strigose or glabrous; taproot often chambered. Shrubs or subshrubs, compact to spreading or rounded and more or less erect, occasionally decumbent, infrequently scapose, (1–)2–15 × 2–25(–30) dm, tomentose to canescent, floccose, or glabrous.
Stems

caudex absent;

aerial flowering stems usually 1, not fistulose, 2–13 dm, strigose or glabrous.

sprawling or spreading to erect, often with persistent leaf bases, up to 1/2 or more height of plant;

caudex stems absent or matted to spreading;

aerial flowering stems erect to spreading, slender, solid, not fistulose, 0.3–2.5(–3) dm, tomentose, canescent, or glabrous.

Leaves

basal and sometimes cauline;

basal petiolate, petiole 2–6 cm, stigose to woolly or glabrous, blade linear-lanceolate or lanceolate to oblanceolate to spatulate, (3–)5–20 × 0.3–2 cm, strigose, becoming glabrous and green on both surfaces except for margins and midvein;

cauline sessile, blade linear-oblanceolate to lanceolate, 1–9 × 0.3–0.8(–1) cm, similar to basal blade.

cauline, 1 per node or fasciculate;

petiole 0.1–0.3 cm, canescent;

blade linear to linear-oblanceolate or oblanceolate, 0.6–1.5(–1.8) × 0.05–0.4(–0.6) cm, white-tomentose or canescent to subglabrous abaxially, tomentose or canescent and grayish, subglabrous, or glabrous and green adaxially, margins often revolute.

Inflorescences

2–10 dm;

branches strigose or glabrous;

bracts semileaflike proximally, linear to linear-lanceolate, 2–9 × 1–3 mm, scalelike distally, triangular, 0.8–5 × 0.5–2 mm.

cymose, infrequently cymose-umbellate or capitate, compact to open, occasionally flat-topped, 0.2–20 × 0.2–15 cm;

branches dichotomous, infrequently absent, tomentose to canescent or glabrous;

bracts usually 3, scalelike, triangular, and 1–3 mm, or leaflike, linear to oblanceolate, and 3–10 × 1–3 mm.

Peduncles

erect, straight or curving upward, 0.5–3.5 cm, strigose or glabrous.

absent.

Involucres

turbinate to campanulate, 2–4(–4.5) × 2–4(–4.5) mm, strigose or glabrous;

teeth 1–1.8 mm.

(1–)3–8 per cluster, turbinate to campanulate, 2–4 × 1.5–3 mm, canescent, pubescent, glabrous, or subglabrous;

teeth 5, erect, 0.3–1.2 mm.

Flowers

1.5–2.5 mm in anthesis, 3–6 mm in fruit;

perianth yellow to yellowish green, rarely maroon in anthesis, often reddish or maroon in fruit, glabrous;

tepals lanceolate;

stamens 1.5–3 mm;

filaments glabrous.

2.5–3 mm;

perianth white to pinkish, glabrous or pubescent;

tepals connate proximal 1/4, monomorphic, usually elliptic to obovate;

stamens exserted, 2.5–5 mm;

filaments subglabrous or pubescent proximally.

Achenes

yellowish green to reddish brown, 5–9 mm, glabrous, 3-winged entire length, beakless.

light brown to brown, 1.8–2.5 mm, glabrous.

Eriogonum alatum

Eriogonum fasciculatum

Distribution
from FNA
AZ; CO; KS; NE; NM; OK; TX; UT; WY; including Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT; including nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The mature reddish roots of Eriogonum alatum can be distinctively chambered. The Navajo (Diné) people consider the species to be a “life medicine” (L. C. Wyman and S. K. Harris 1951), using a mixture of shredded roots and water primarily to treat internal ailments. The species is used also as a ceremonial medicine (P. A. Vestal 1952). The Zuni use it as an emetic for stomachaches (S. Camazine and R. A. Bye 1980).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 5 (4 in the flora).

Eriogonum fasciculatum is a complex, polyploid series of variants that are generally distinct but often difficult to distinguish morphologically. Variety emphereium Reveal is confined to central Baja California, Mexico. The introduction of Eriogonum fasciculatum as a decorative roadside plant by the California Department of Transportation is resulting in hybrid populations involving E. cinereum. The aggressively weedy and (for Arizona) exotic variety foliosum is rapidly invading the native habitat of var. polifolium. Members of E. fasciculatum are food plants for several butterflies, notably the Bernardino dotted-blue (Euphilotes bernardino), lupine blue (Plebeius lupini), Mormon metalmark (Apodemia mormo), and Behr’s metalmark (A. virgulti). Probably the butterfly most commonly seen with the species is the nut-brown hairstreak (Satyrium saepium), which frequents plants in full flower. Eriogonum fasciculatum is also the most important native source of honey in California.

This widespread species was used extensively by Native Americans for a variety of ailments. Its application for pain and headaches (D. P. Barrows 1900; K. Hedges 1986; E. W. Voegelin 1938) was rather common, as was its general use for diarrhea (Hedges; Voegelin). M. L. Zigmond (1981) reported that the Kawaiisu lined their acorn granaries with leaves of var. proliferum to keep out rain—a daunting challenge given the size of the leaves! L. Hinton (1975) reported the use of a decoction of dried flowers and roots to maintain a healthy heart, and M. C. Stevenson (1915) indicated that a powder derived from the roots was used by the Zuñi to treat wounds, whereas a root decoction was taken for colds and hoarseness. B. R. Bocek (1984) reported that the Costanoan Indians of California used a decoction of the plant to treat unspecified urinary problems. F. H. Elmore (1943) reported the use of a decoction of var. proliferum by the Navajo (Diné) people as an anti-witchcraft medicine.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowering stems and inflorescence branches usually strigose; peduncles strigose or nearly so; involucres strigose, rarely glabrous; plants 5-13(-17) dm; n Arizona, Colorado, w Kansas, sw Nebraska, New Mexico, w Texas, e Utah, se Wyoming
var. alatum
1. Flowering stems and inflorescence branches glabrous or nearly so; peduncles glabrous, occasionally slightly strigose; involucres glabrous; plants 10-20(-25) dm; ne New Mexico, w Oklahoma, n Texas
var. glabriusculum
1. Leaves thinly white-tomentose abaxially, glabrous adaxially; perianths and involucres mostly glabrous; plants usually decumbent; primarily coastal 80c. Eriogonum fasciculatum var. fasciculatum
→ 1
1. Leaves, perianths, and involucres pubescent or, if subglabrous, then plants of desert regions; plants erect to rounded; generally inland
→ 2
2. Leaves light yellow-green, subglabrous adaxially; involucres and perianths glabrous or subglabrous
var. flavoviride
2. Leaves dark green or grayish, usually pubescent adaxially; involucres and perianths pubescent
→ 3
3. Inflorescences capitate to cymose-umbellate, rarely cymose; leaves canescent on both surfaces or densely tomentose abaxially and canescent adaxially; leaf margins plane or infrequently revolute; mainly deserts
var. polifolium
3. Inflorescences open and mostly cymose; leaves densely white-tomentose abaxially, less so to green and floccose adaxially; leaf margins usually tightly revolute; mainly inner coastal foothills
var. foliolosum
Source FNA vol. 5, p. 379. FNA vol. 5, p. 297.
Parent taxa Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Pterogonum Polygonaceae > subfam. Eriogonoideae > Eriogonum > subg. Eucycla
Sibling taxa
E. abertianum, E. acaule, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. fasciculatum, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
E. abertianum, E. acaule, E. alatum, E. aliquantum, E. allenii, E. alpinum, E. ammophilum, E. ampullaceum, E. androsaceum, E. anemophilum, E. angulosum, E. annuum, E. apiculatum, E. apricum, E. arborescens, E. arcuatum, E. aretioides, E. argillosum, E. argophyllum, E. arizonicum, E. artificis, E. atrorubens, E. baileyi, E. batemanii, E. bicolor, E. bifurcatum, E. brachyanthum, E. brachypodum, E. brandegeei, E. breedlovei, E. brevicaule, E. butterworthianum, E. caespitosum, E. capillare, E. cernuum, E. chrysops, E. cinereum, E. cithariforme, E. clavatum, E. clavellatum, E. codium, E. collinum, E. coloradense, E. compositum, E. concinnum, E. congdonii, E. contiguum, E. contortum, E. correllii, E. corymbosum, E. covilleanum, E. crocatum, E. cronquistii, E. crosbyae, E. cusickii, E. darrovii, E. dasyanthemum, E. davidsonii, E. deflexum, E. deserticola, E. desertorum, E. diatomaceum, E. diclinum, E. divaricatum, E. douglasii, E. eastwoodianum, E. effusum, E. elatum, E. elegans, E. elongatum, E. ephedroides, E. eremicola, E. eremicum, E. ericifolium, E. esmeraldense, E. evanidum, E. exaltatum, E. exilifolium, E. flavum, E. fusiforme, E. giganteum, E. gilmanii, E. glandulosum, E. gordonii, E. gossypinum, E. gracile, E. gracilipes, E. gracillimum, E. grande, E. greggii, E. gypsophilum, E. havardii, E. heermannii, E. helichrysoides, E. hemipterum, E. heracleoides, E. hieracifolium, E. hirtellum, E. hirtiflorum, E. hoffmannii, E. holmgrenii, E. hookeri, E. howellianum, E. hylophilum, E. incanum, E. inerme, E. inflatum, E. intrafractum, E. jamesii, E. jonesii, E. kelloggii, E. kennedyi, E. kingii, E. lachnogynum, E. lancifolium, E. latens, E. latifolium, E. lemmonii, E. leptocladon, E. leptophyllum, E. libertini, E. lobbii, E. loganum, E. lonchophyllum, E. longifolium, E. luteolum, E. maculatum, E. mancum, E. marifolium, E. mensicola, E. microthecum, E. mitophyllum, E. mohavense, E. molestum, E. mortonianum, E. multiflorum, E. natum, E. nealleyi, E. nervulosum, E. nidularium, E. niveum, E. nortonii, E. novonudum, E. nudum, E. nummulare, E. nutans, E. ochrocephalum, E. ordii, E. ostlundii, E. ovalifolium, E. palmerianum, E. panamintense, E. panguicense, E. parishii, E. parvifolium, E. pauciflorum, E. pelinophilum, E. pendulum, E. pharnaceoides, E. plumatella, E. polycladon, E. polypodum, E. prattenianum, E. prociduum, E. pulchrum, E. pusillum, E. pyrolifolium, E. racemosum, E. reniforme, E. ripleyi, E. rixfordii, E. robustum, E. rosense, E. roseum, E. rotundifolium, E. rubricaule, E. rupinum, E. salicornioides, E. saxatile, E. scabrellum, E. scopulorum, E. shockleyi, E. siskiyouense, E. smithii, E. soliceps, E. soredium, E. spathulatum, E. spectabile, E. spergulinum, E. sphaerocephalum, E. strictum, E. subreniforme, E. suffruticosum, E. temblorense, E. tenellum, E. ternatum, E. terrenatum, E. thomasii, E. thompsoniae, E. thornei, E. thurberi, E. thymoides, E. tiehmii, E. tomentosum, E. trichopes, E. tripodum, E. truncatum, E. tumulosum, E. twisselmannii, E. umbellatum, E. ursinum, E. vestitum, E. villiflorum, E. vimineum, E. viridescens, E. viridulum, E. viscidulum, E. visheri, E. watsonii, E. wetherillii, E. wootonii, E. wrightii, E. zionis
Subordinate taxa
E. alatum var. alatum, E. alatum var. glabriusculum
E. fasciculatum var. flavoviride, E. fasciculatum var. foliolosum, E. fasciculatum var. polifolium
Synonyms Pterogonum alatum
Name authority Torrey: in L. Sitgreaves, Rep. Exped. Zuni Colorado Rivers, 168, plate 8. (1853) Bentham: Trans. Linn. Soc. London 17: 411. (1836)
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