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flattened pipewort

pipewort family

Habit Herbs, perennial, 20–70 cm. Herbs, annual or perennial, cespitose or solitary, rosulate, rarely caulescent, mostly scapose, glabrous or variously pubescent with simple or compound hairs.
Stems

erect to repent or prostrate, axis sympodial or monopodial, sometimes branching.

Leaves

linear-attenuate, 5–30 cm, apex subulate.

mostly many ranked in rosettes, sometimes in loose spirals, mostly grasslike;

blade linear to linear-triangular, lingulate, tapering, base mostly clasping;

veins parallel.

Inflorescences

scape sheaths mostly longer than principal leaves, loose;

scapes linear, 1–3 mm wide, multiribbed (ribs lacunar);

heads chalk white except for dark gray or near black exserted tips of receptacular bracts, anthers, hemispheric to subglobose, 10–20 mm wide, soft, much flattened when pressed;

receptacle pilose;

involucral bracts frequently squarrose, later obscured by mature bracteoles and flowers, gray, broadly ovate to oblong or elliptic, 2–3 mm, margins entire, apex rounded or obtuse, glabrous;

inner bracts, receptacular bracteoles dark gray, spatulate-linear to oblong, 2–3 mm, margins entire, apex acute with white, club-shaped hairs.

terminal and/or axillary, simple [compound], involucrate [proliferative], buttonlike or short-cylindric;

scape sheaths spathelike, tubular, enclosing scape base, distally open;

scapes 1–many, simple [compound], terete, usually twisted, mostly multiribbed;

involucral bracts spirally imbricate series, usually chaffy or scarious, grading inward to receptacular bracts or these absent;

receptacle glabrous or variously hairy.

Flowers

(florets) unisexual, staminate and pistillate on same [rarely different] plants, radially or bilaterally symmetric;

sepals 2–3, distinct or variously connate, diverging from stipelike base or merely part of lobeless column;

petals 0 or 2–3, diverging from short to elongate stipe (part of androphore or gynophore) or merely part of lobeless column;

stamens 2–4(–6), often unequal;

filaments arising from floral axis, rarely epipetalous;

anthers mostly black, 1–2-locular, 2–4-sporangiate, versatile or basifixed, dehiscence longitudinal;

pollen 1-grooved, 3-nucleate;

appendages present in some flowers, glandlike or peglike, rarely bladelike, sometimes fringed or variously cleft;

pistil compound, 2–3-carpellate;

ovary superior, raised on gynophore, 1 locule per carpel;

ovules 1 per locule, adaxial-apical, pendulous, orthotropous, bitegmic;

style terminal, appendaged or unappendaged, 2–3-branched, branches simple or apex rebranched.

Staminate flowers

sepals 2, pale or with dark apex, linear or linear-spatulate, 2–4 mm, apex acute to blunt with mealy white, club-shaped hairs;

androphore broadly club-shaped;

petals 2, pale, oblong, conspicuously unequal, larger lobe apically fringed with pale, club-shaped hairs, smaller lobe glabrous or with a few hairs at apex;

stamens 3–4(–6);

anthers black.

Pistillate flowers

sepals 2, dark at apex, oblong-spatulate, 2.5–3 mm, abaxially with mealy white, club-shaped hairs, adaxially with translucent hairs;

petals 2, pale, oblong-spatulate, apex acute, abaxially with mealy white, club-shaped hairs, adaxially with translucent hairs;

pistil 2-carpellate.

Fruits

capsules, thin-walled, loculicidal.

Seeds

dark lustrous brown, broadly ovoid to near round but asymmetric, 0.5 mm, mostly minutely spiny-papillate.

translucent, ovoid, ellipsoid, or broadly fusiform, mostly 1 mm or shorter, variously ribbed or sculptured;

endosperm copious, mealy-starchy, with compound starch grains;

embryo apical.

Rootstocks

rhizomes or stems, thickened, short to variously elongate;

roots fibrous, spongy, or spongy-septate (diaphragmatic).

Eriocaulon compressum

Eriocaulaceae

Phenology Flowering late winter–spring.
Habitat Sands and peats of shallow pineland ponds, savanna, seeps, ditches, or low flatwoods
Elevation 0–300 m (0–1000 ft)
Distribution
from FNA
AL; DE; FL; GA; LA; MD; MS; NC; NJ; SC; TX
[WildflowerSearch map]
[BONAP county map]
Nearly worldwide; primarily tropical and subtropical regions
[BONAP county map]
Discussion

Eriocaulon compressum is polymorphic in habit. Male flowers vary considerably in length and degree of connation, and female flowers are often sterile. Of the southeastern coastal plain species this and the similar, but proportionately smaller, E. lineare, are the most aquatic, the former most common around clay-based ponds, the latter around karst ponds. Unlike the more northern E. aquaticum, these two species seldom frequent the shallows of flowing streams.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Most Eriocaulaceae grow in full sun in acidic wet soils or in aquatic situations.

Genera ca. 13, species 1200+ (3 genera, 16 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lacunar tissue evident at leaf base; larger roots pale, thickened, septate, unbranched; perianth parts in 2s (except in Eriocaulon cinereum and E. microcephalum), petals with glands on adaxial surface; stamens 3–4 or 6; anthers 2-locular, apex of staminal column with 2–3 glands, glands unappendaged; pistil 2(–3)-carpellate; style unappendaged, 2(–3)-branched
Eriocaulon
1. Lacunar tissue not evident at leaf base; larger roots either dark, fibrous, and evidently branched or pale, thickened, and spongy, neither septate nor branched; perianth parts in 3s (except in Lachnocaulon digynum), petals if present without glands; stamens (2–)3; anthers 1–2-locular; apex of staminal column usually with 3 glands, glands appendaged or unappendaged; pistil (2–)3-carpellate; style appendaged, 2–3-branched.
→ 2
2. Roots dark, slender, fibrous, evidently branched; scapes glabrous or hairy, hairs neither swollen nor glandular; expanded inflorescences ovoid to globose or short-cylindric, basal involucral bracts reflexed, obscured by inflorescence; hairs of perianth club-shaped; staminal filaments adnate to rim of androphore; anthers 1-locular; style branches 2-cleft
Lachnocaulon
2. Roots pale, thickened, spongy, appearing unbranched; scapes hairy, at least some hairs swollen basally, often glandular distally; expanded inflorescences hemispheric to globose, basal bracts not obscured by inflorescence; hairs of perianth tapering, acute, not club-shaped; staminal filaments low in corolla tube; anthers 2-locular; style branches undivided
Syngonanthus
Source FNA vol. 22. FNA vol. 22, p. 198. Author: Robert Kral.
Parent taxa Eriocaulaceae > Eriocaulon
Sibling taxa
E. aquaticum, E. cinereum, E. decangulare, E. koernickianum, E. lineare, E. microcephalum, E. nigrobracteatum, E. parkeri, E. ravenelii, E. texense
Subordinate taxa
Eriocaulon, Lachnocaulon, Syngonanthus
Synonyms E. decangulare, E. gnaphalodes, E. cephalotes, Sphaerochloa compressa
Name authority Lamarck: in J. Lamarck et al., Encycl. 3: 276. (1789) Palisot de Beauvois ex Desvaux
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