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shaggy daisy, shaggy fleabane, tall annual willowherb

large-flower daisy, large-flower fleabane, onestem fleabane, Rocky Mountain alpine fleabane

Habit Perennials, 5–30(–50) cm; taprooted, caudices with relatively short and thick branches. Perennials, 2–25 cm; rhizomatous, fibrous-rooted, caudices or rhizomes crownlike or branches relatively short and thick.
Stems

erect, hirsute to hispido-hirsute (often with slightly deflexed hairs), minutely to stipitate glandular.

erect to decumbent-ascending, sparsely to moderately pilose to villoso-hirsute, often stipitate-glandular over all or part.

Leaves

basal (persistent) and cauline (petioles prominently ciliate, hairs thick-based, spreading);

basal blades oblanceolate to narrowly oblanceolate, 20–80 × 1–4(–5) mm, margins entire, faces hispid to hispido-hirsute, little, if at all, glandular;

cauline on distal 1/2–3/4 of stems, blades becoming linear-lanceolate, little reduced distally.

basal (persistent) and cauline (petioles equaling or shorter than blades);

blades oblanceolate to obovate or spatulate, 10–60(–90) × 3–8(–14) mm, cauline abruptly or gradually reduced distally, margins entire (apices rounded), faces sparsely hirsutulous or villous to sparsely strigose or glabrate, sometimes sparsely glandular.

Involucres

4–7 × 7–15 mm.

5–8(–10) × 8–20 mm.

Ray florets

50–100;

corollas white to pink, less commonly bluish, 6–15 mm, laminae reflexing.

50–130;

corollas blue to pink or purplish, rarely white, 7–11(–15) mm (mostly 1–2 mm wide), laminae coiling.

Disc corollas

3–5 mm (throats distinctly indurate and inflated, glabrous or sparsely puberulent with glandular-viscid, hairs blunt).

2.4–4(–5) mm.

Phyllaries

in 2–4 series (midvein region orange to yellowish), hirsute to hispido-hirsute, minutely glandular.

in 2–3 series (green or purplish), moderately to densely woolly-villous (hairs flattened, cross walls sometimes reddish), minutely glandular at least apically.

Heads

1–5(–50).

1.

Cypselae

1.4–1.8 mm, 2-nerved, faces sparsely strigose;

pappi: outer of setae or subulate scales (0.1–0.3 mm), inner of 12–27 bristles.

1.8–2.4 mm, 2-nerved, faces strigose;

pappi: outer of setae, inner of (7–)10–18(–22) bristles.

2n

= 18, 27.

Erigeron pumilus

Erigeron grandiflorus

Phenology Flowering Jul–Aug(–Sep).
Habitat Rocky sites, meadows, alpine or near timberline
Elevation 2900–4200 m (9500–13800 ft)
Distribution
from FNA
CA; CO; ID; KS; MT; ND; NE; NV; OR; SD; UT; WA; WY; AB; BC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; ID; MT; NM; OR; UT; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

S. A. Spongberg (1971) recognized only the triploid populations as Erigeron grandiflorus and assigned the diploid ones to E. simplex. He hypothesized that the triploids incorporate genomic elements from an ancestor other than E. simplex. Based on his comments and annotations, however, triploids in southern Canada and the western United States apparently differ from the much more widespread diploids only quantitatively, having involucres and florets at the higher end of size ranges. Morphologic distinctions between the ploidal races do not provide a basis for consistent distinction. Spongberg (p. 200) also noted that “because of the intergrading of morphologic features of plants of Erigeron grandiflorus...the single most important criterion indicative of this taxon is highly irregular [in shape] and greatly abortive pollen.” These pollen features result from meiotic anomalies associated with the triploid condition.

Specimen citations by A. Cronquist (1947) for Erigeron grandiflorus were mostly from collections of the species treated here as E. porsildii. He also cited two collections from southwestern Alberta; those and the type collection of E. grandiflorus (from the same region) are disjunct by more than 1500 kilometers from the more northern range of E. porsildii and instead lie at the northern extremity of the range of what previously has generally been identified as E. simplex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pappi: outer of coarse bristles or setiform scales, inner of 12–20 bristles; rays usually pink, sometimes white; indurate portion of disc corollas relatively dull, glabrous or slightly puberulent
var. intermedius
1. Pappi: outer of inconspicuous, well-developed setae, inner of 15–27 bristles; rays white; indurate portion of disc corollas shiny, glabrous
var. pumilus
Source FNA vol. 20, p. 288. FNA vol. 20, p. 324.
Parent taxa Asteraceae > tribe Astereae > Erigeron Asteraceae > tribe Astereae > Erigeron
Sibling taxa
E. abajoensis, E. acomanus, E. acris, E. aequifolius, E. algidus, E. aliceae, E. allocotus, E. alpiniformis, E. anchana, E. annuus, E. aphanactis, E. arenarioides, E. argentatus, E. arisolius, E. arizonicus, E. asperugineus, E. aureus, E. barbellulatus, E. basalticus, E. bellidiastrum, E. bigelovii, E. biolettii, E. blochmaniae, E. bloomeri, E. breweri, E. caespitosus, E. calvus, E. canaani, E. canus, E. cascadensis, E. cavernensis, E. cervinus, E. chrysopsidis, E. clokeyi, E. compactus, E. compositus, E. concinnus, E. consimilis, E. corymbosus, E. coulteri, E. cronquistii, E. davisii, E. decumbens, E. denalii, E. disparipilus, E. divergens, E. eatonii, E. elatior, E. elatus, E. elegantulus, E. elmeri, E. engelmannii, E. evermannii, E. eximius, E. filifolius, E. flabellifolius, E. flagellaris, E. flettii, E. foliosus, E. formosissimus, E. garrettii, E. geiseri, E. glabellus, E. glacialis, E. glaucus, E. goodrichii, E. gracilis, E. grandiflorus, E. greenei, E. heliographis, E. hessii, E. howellii, E. humilis, E. hyperboreus, E. hyssopifolius, E. inornatus, E. jonesii, E. kachinensis, E. karvinskianus, E. klamathensis, E. kuschei, E. lackschewitzii, E. lanatus, E. lassenianus, E. latus, E. leibergii, E. leiomerus, E. lemmonii, E. linearis, E. lobatus, E. lonchophyllus, E. maguirei, E. mancus, E. maniopotamicus, E. mariposanus, E. melanocephalus, E. miser, E. modestus, E. muirii, E. multiceps, E. nanus, E. nauseosus, E. nematophyllus, E. neomexicanus, E. nivalis, E. ochroleucus, E. oreganus, E. oreophilus, E. ovinus, E. oxyphyllus, E. pallens, E. parishii, E. parryi, E. peregrinus, E. petrophilus, E. philadelphicus, E. pinnatisectus, E. piperianus, E. piscaticus, E. poliospermus, E. porsildii, E. pringlei, E. procumbens, E. pulchellus, E. pulcherrimus, E. purpuratus, E. pygmaeus, E. quercifolius, E. radicatus, E. reductus, E. religiosus, E. rhizomatus, E. robustior, E. rybius, E. rydbergii, E. salishii, E. salmonensis, E. sanctarum, E. saxatilis, E. sceptrifer, E. scopulinus, E. serpentinus, E. sionis, E. sivinskii, E. sparsifolius, E. speciosus, E. strigosus, E. subglaber, E. subtrinervis, E. supplex, E. tenellus, E. tener, E. tenuis, E. tracyi, E. trifidus, E. tweedyi, E. uintahensis, E. uncialis, E. uniflorus, E. untermannii, E. ursinus, E. utahensis, E. vagus, E. velutipes, E. vernus, E. versicolor, E. vetensis, E. vicinus, E. vreelandii, E. watsonii, E. wilkenii, E. yukonensis
E. abajoensis, E. acomanus, E. acris, E. aequifolius, E. algidus, E. aliceae, E. allocotus, E. alpiniformis, E. anchana, E. annuus, E. aphanactis, E. arenarioides, E. argentatus, E. arisolius, E. arizonicus, E. asperugineus, E. aureus, E. barbellulatus, E. basalticus, E. bellidiastrum, E. bigelovii, E. biolettii, E. blochmaniae, E. bloomeri, E. breweri, E. caespitosus, E. calvus, E. canaani, E. canus, E. cascadensis, E. cavernensis, E. cervinus, E. chrysopsidis, E. clokeyi, E. compactus, E. compositus, E. concinnus, E. consimilis, E. corymbosus, E. coulteri, E. cronquistii, E. davisii, E. decumbens, E. denalii, E. disparipilus, E. divergens, E. eatonii, E. elatior, E. elatus, E. elegantulus, E. elmeri, E. engelmannii, E. evermannii, E. eximius, E. filifolius, E. flabellifolius, E. flagellaris, E. flettii, E. foliosus, E. formosissimus, E. garrettii, E. geiseri, E. glabellus, E. glacialis, E. glaucus, E. goodrichii, E. gracilis, E. greenei, E. heliographis, E. hessii, E. howellii, E. humilis, E. hyperboreus, E. hyssopifolius, E. inornatus, E. jonesii, E. kachinensis, E. karvinskianus, E. klamathensis, E. kuschei, E. lackschewitzii, E. lanatus, E. lassenianus, E. latus, E. leibergii, E. leiomerus, E. lemmonii, E. linearis, E. lobatus, E. lonchophyllus, E. maguirei, E. mancus, E. maniopotamicus, E. mariposanus, E. melanocephalus, E. miser, E. modestus, E. muirii, E. multiceps, E. nanus, E. nauseosus, E. nematophyllus, E. neomexicanus, E. nivalis, E. ochroleucus, E. oreganus, E. oreophilus, E. ovinus, E. oxyphyllus, E. pallens, E. parishii, E. parryi, E. peregrinus, E. petrophilus, E. philadelphicus, E. pinnatisectus, E. piperianus, E. piscaticus, E. poliospermus, E. porsildii, E. pringlei, E. procumbens, E. pulchellus, E. pulcherrimus, E. pumilus, E. purpuratus, E. pygmaeus, E. quercifolius, E. radicatus, E. reductus, E. religiosus, E. rhizomatus, E. robustior, E. rybius, E. rydbergii, E. salishii, E. salmonensis, E. sanctarum, E. saxatilis, E. sceptrifer, E. scopulinus, E. serpentinus, E. sionis, E. sivinskii, E. sparsifolius, E. speciosus, E. strigosus, E. subglaber, E. subtrinervis, E. supplex, E. tenellus, E. tener, E. tenuis, E. tracyi, E. trifidus, E. tweedyi, E. uintahensis, E. uncialis, E. uniflorus, E. untermannii, E. ursinus, E. utahensis, E. vagus, E. velutipes, E. vernus, E. versicolor, E. vetensis, E. vicinus, E. vreelandii, E. watsonii, E. wilkenii, E. yukonensis
Subordinate taxa
E. pumilus var. intermedius, E. pumilus var. pumilus
Synonyms E. simplex
Name authority Nuttall: Gen. N. Amer. Pl. 2: 147. (1818) Hooker: Fl. Bor.-Amer. 2: 18, plate 123. (1834)
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