Erigeron grandiflorus |
Erigeron tracyi |
|
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large-flower daisy, large-flower fleabane, onestem fleabane, Rocky Mountain alpine fleabane |
running fleabane |
|
Habit | Perennials, 2–25 cm; rhizomatous, fibrous-rooted, caudices or rhizomes crownlike or branches relatively short and thick. | Annuals, biennials, or short-lived perennials, 2.5–8(–12, 18) cm; usually taprooted, sometimes fibrous-rooted, caudices simple or branched. |
Stems | erect to decumbent-ascending, sparsely to moderately pilose to villoso-hirsute, often stipitate-glandular over all or part. |
first erect (greenish proximally), then producing herbaceous, leafy, prostrate runners (stoloniform branches, sometimes with rooting plantlets at tips), densely hirsutulous (hairs spreading-deflexed, of relatively even lengths and orientations), sparsely minutely glandular. |
Leaves | basal (persistent) and cauline (petioles equaling or shorter than blades); blades oblanceolate to obovate or spatulate, 10–60(–90) × 3–8(–14) mm, cauline abruptly or gradually reduced distally, margins entire (apices rounded), faces sparsely hirsutulous or villous to sparsely strigose or glabrate, sometimes sparsely glandular. |
mostly basal (persistent in early season); blades oblanceolate to spatulate (obovate-elliptic laminae), 10–30(–60) × 3–6(–12) mm, cauline abruptly reduced distally, margins entire, dentate, or lobed, faces densely hirsute, eglandular. |
Involucres | 5–8(–10) × 8–20 mm. |
3.5–4.5(–6) × 6–9(–12) mm. |
Ray florets | 50–130; corollas blue to pink or purplish, rarely white, 7–11(–15) mm (mostly 1–2 mm wide), laminae coiling. |
60–130; corollas white, often purplish abaxially, sometimes with an abaxial midstripe, 5–9 mm, laminae not coiling or reflexing. |
Disc corollas | 2.4–4(–5) mm. |
2–3 mm (throats indurate and slightly inflated). |
Phyllaries | in 2–3 series (green or purplish), moderately to densely woolly-villous (hairs flattened, cross walls sometimes reddish), minutely glandular at least apically. |
in 3–4 series, sparsely to moderately hirsute, minutely glandular. |
Heads | 1. |
1(–3 rarely, from midstem or proximal branches). |
Cypselae | 1.8–2.4 mm, 2-nerved, faces strigose; pappi: outer of setae, inner of (7–)10–18(–22) bristles. |
0.7–1.3 mm, 2-nerved, faces sparsely strigose; pappi: outer of setae, inner of 12–16 bristles. |
2n | = 18, 27. |
= 27. |
Erigeron grandiflorus |
Erigeron tracyi |
|
Phenology | Flowering Jul–Aug(–Sep). | Flowering Mar–Oct. |
Habitat | Rocky sites, meadows, alpine or near timberline | Desert scrub, grassy slopes, oak chaparral, pinyon-juniper woodlands, Douglas fir-ponderosa pine |
Elevation | 2900–4200 m (9500–13800 ft) | 700–2300(–2400) m (2300–7500(–7900) ft) |
Distribution |
AZ; CO; ID; MT; NM; OR; UT; WY; AB; BC
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AZ; CO; KS; NM; NV; OK; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Durango, Sonora, Zacatecas)
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Discussion | S. A. Spongberg (1971) recognized only the triploid populations as Erigeron grandiflorus and assigned the diploid ones to E. simplex. He hypothesized that the triploids incorporate genomic elements from an ancestor other than E. simplex. Based on his comments and annotations, however, triploids in southern Canada and the western United States apparently differ from the much more widespread diploids only quantitatively, having involucres and florets at the higher end of size ranges. Morphologic distinctions between the ploidal races do not provide a basis for consistent distinction. Spongberg (p. 200) also noted that “because of the intergrading of morphologic features of plants of Erigeron grandiflorus...the single most important criterion indicative of this taxon is highly irregular [in shape] and greatly abortive pollen.” These pollen features result from meiotic anomalies associated with the triploid condition. Specimen citations by A. Cronquist (1947) for Erigeron grandiflorus were mostly from collections of the species treated here as E. porsildii. He also cited two collections from southwestern Alberta; those and the type collection of E. grandiflorus (from the same region) are disjunct by more than 1500 kilometers from the more northern range of E. porsildii and instead lie at the northern extremity of the range of what previously has generally been identified as E. simplex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In March through June, plants of Erigeron tracyi produce leaves in a basal rosette usually with a single, erect, monocephalous, stem. Stoloniform branches are soon formed (often recognized on pressed specimens by the leaves mostly on one side of the branches), and by the end of the season (August through October), prostrate runners are usually evident, sometimes forming terminal, rooting plantlets. Erigeron tracyi is similar in habit to E. flagellaris, particularly in the herbaceous stolons or stoloniform branches; the stem pubescence of E. tracyi is different, the stolons much less commonly produce rooting plantlets at the tips, and the plants tend to be perennial with woody or lignescent caudices, although they are variable both in habit and duration. Apparent hybrids with E. modestus and E. flagellaris are occasionally encountered, and the most common form of E. tracyi is perhaps (speculative) a stabilized, apomictic hybrid between the latter and E. divergens. All chromosome counts thus far have shown E. tracyi to be triploid and asynaptic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 324. | FNA vol. 20, p. 340. |
Parent taxa | Asteraceae > tribe Astereae > Erigeron | Asteraceae > tribe Astereae > Erigeron |
Sibling taxa | ||
Synonyms | E. simplex | E. cinereus, E. colomexicanus, E. commixtus, E. divergens var. cinereus |
Name authority | Hooker: Fl. Bor.-Amer. 2: 18, plate 123. (1834) | Greene: Pittonia 5: 59. (1902) |
Web links |