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King's sandwort

Photo is of parent taxon

King's sandwort, King's smooth sandwort

Habit Plants tufted or sometimes in compact cushion, green, not glaucous, woody or not at base.
Stems

erect, (1–)3–20(–25) cm, stipitate-glandular or glabrous proximally.

Leaves

basal leaves abundant, persistent;

cauline leaves in (1–)4+ pairs, reduced distally or not;

basal blades erect or closely ascending to somewhat spreading, green to gray-green, filiform to needlelike or narrowly subulate, 0.3–3(–4) cm × 0.3–1.2 mm, flexuous or rigid, herbaceous, apex apiculate or stiff and spinose, glabrous to stipitate-glandular, not glaucous.

Inflorescences

(1–)3–13-flowered, ± open cymes.

Pedicels

2–15 mm, glabrous to densely stipitate-glandular.

Flowers

sepals 1–3-veined, lateral veins less developed, ovate or lanceolate, (2.5–)2.8–5(–6) mm, not expanding in fruit, margins broad, apex obtuse to broadly acute or acuminate, glabrous or stipitate-glandular on herbaceous portion;

petals white or rarely pink, oblong to spatulate, (3–)4–7 mm, ca. 1.2–1.3 times as long as sepals, apex entire, erose, or 2-fid almost to base;

nectaries as abaxial, rounded lobe with transverse groove or elongate cup at base of filaments opposite sepals, 0.6 × 0.3 mm.

Capsules

4.5–7 mm, glabrous.

Seeds

black to brown, spheric or oblong to ovoid, 1.2–2.1 mm, low-tuberculate, sometimes papillate on abaxial ridge.

Petals

white (rarely pink), apex entire or erose.

Eremogone kingii

Eremogone kingii var. glabrescens

Phenology Flowering late spring–summer.
Habitat Dry hills, pinyon-juniper woods, sagebrush slopes
Elevation (1500-)2100-4000 m [(4900-)6900-13100 ft]
Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR; UT
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

M. F. Baad (1969) considered Eremogone kingii to be monophyletic despite considerable morphological variation; J. C. Hickman (1971) thought otherwise, considering E. kingii to be a “genetic dumping ground for all the closely related taxa,” but did not propose any new taxonomic alignment.

Eremogone kingii is extremely variable throughout its range with six infraspecific taxa recognized (under Arenaria kingii) by B. Maguire (1947, 1951). We have been unsuccessful in distinguishing more than two of those taxa. The others intergrade to such an extent that formal recognition is unwarranted. Most distinctive of these here-rejected taxa is var. uintahensis, said to have sepals (4.5–)5–6 mm, versus 3.6–4.5(–5) for the other taxa. Interestingly, the type specimen has sepals mostly 4.5 mm long. In the main portion of the range of var. uintahensis, the sepals are rounded to broadly obtuse, but they may also be acute. Furthermore, the sepals and pedicels are often glabrous, but the correlation of the above-mentioned characters varies over the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Arenaria kingii subsp. compacta intergrades completely with var. glabrescens in many areas of the Sierra Nevada and does not deserve recognition.

M. F. Baad (1969) noted populations from Idaho, Nevada, Oregon, and Wyoming that he considered intermediate between Eremogone capillaris var. americana and E. kingii var. glabrescens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Caryophyllaceae > subfam. Alsinoideae > Eremogone Caryophyllaceae > subfam. Alsinoideae > Eremogone > Eremogone kingii
Sibling taxa
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. franklinii, E. hookeri, E. macradenia, E. pumicola, E. stenomeres, E. ursina
E. kingii var. kingii
Subordinate taxa
E. kingii var. glabrescens, E. kingii var. kingii
Key
1. Petals white, apex 2-fid
var. kingii
1. Petals white or rarely pink, apex entire or erose
var. glabrescens
Synonyms Stellaria kingii, Arenaria kingii Arenaria fendleri var. glabrescens, Arenaria aculeata var. uintahensis, Arenaria compacta, Arenaria kingii subsp. compacta, Arenaria kingii var. glabrescens, Arenaria kingii subsp. plateauensis, Arenaria kingii var. plateauensis, Arenaria kingii subsp. rosea, Arenaria kingii subsp. uintahensis, Arenaria kingii var. uintahensis, Arenaria uintahensis, E. kingii var. plateauensis, E. kingii var. rosea, E. kingii subsp. uintahensis
Name authority (S. Watson) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 140. (1973) (S. Watson) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001)
Source FNA vol. 5, p. 67. Treatment authors: Ronald L. Hartman, Richard K. Rabeler, Frederick H. Utech. FNA vol. 5, p. 68. Treatment authors: Ronald L. Hartman, Richard K. Rabeler, Frederick H. Utech.
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