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King's sandwort

Eremogone franklinii

Franklin's sandwort

Habit Plants tufted or sometimes in compact cushion, green, not glaucous, woody or not at base. Plants ± cespitose, bluish green, not glaucous, with woody base.
Stems

erect, (1–)3–20(–25) cm, stipitate-glandular or glabrous proximally.

erect, 3–10(–15) cm, glabrous.

Leaves

basal leaves abundant, persistent;

cauline leaves in (1–)4+ pairs, reduced distally or not;

basal blades erect or closely ascending to somewhat spreading, green to gray-green, filiform to needlelike or narrowly subulate, 0.3–3(–4) cm × 0.3–1.2 mm, flexuous or rigid, herbaceous, apex apiculate or stiff and spinose, glabrous to stipitate-glandular, not glaucous.

basal leaves persistent;

cauline leaves in 6–10 pairs, closely overlapping, not reduced;

basal blades arcuate-spreading, needlelike, (0.6–)1–2 cm × ca. 1 mm, ± rigid, herbaceous, apex spinose, glabrous, not glaucous.

Inflorescences

(1–)3–13-flowered, ± open cymes.

3–45+-flowered, usually congested, subcapitate cymes.

Pedicels

2–15 mm, glabrous to densely stipitate-glandular.

ca. 0.1–3(–4) mm, glabrous.

Flowers

sepals 1–3-veined, lateral veins less developed, ovate or lanceolate, (2.5–)2.8–5(–6) mm, not expanding in fruit, margins broad, apex obtuse to broadly acute or acuminate, glabrous or stipitate-glandular on herbaceous portion;

petals white or rarely pink, oblong to spatulate, (3–)4–7 mm, ca. 1.2–1.3 times as long as sepals, apex entire, erose, or 2-fid almost to base;

nectaries as abaxial, rounded lobe with transverse groove or elongate cup at base of filaments opposite sepals, 0.6 × 0.3 mm.

sepals 1–3-veined, linear-lanceolate, (5–)8.5–12 mm, not enlarging in fruit, margins narrow, apex acuminate, glabrous;

petals white, oblong-lanceolate to narrowly spatulate, 7–9 mm, 0.8–1.1 times as long as sepals, apex rounded to blunt;

nectaries not apparent.

Capsules

4.5–7 mm, glabrous.

2.3–3.3 mm, glabrous.

Seeds

black to brown, spheric or oblong to ovoid, 1.2–2.1 mm, low-tuberculate, sometimes papillate on abaxial ridge.

black, pyriform, 1.2–1.7 mm, tuberculate.

Eremogone kingii

Eremogone franklinii

Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

M. F. Baad (1969) considered Eremogone kingii to be monophyletic despite considerable morphological variation; J. C. Hickman (1971) thought otherwise, considering E. kingii to be a “genetic dumping ground for all the closely related taxa,” but did not propose any new taxonomic alignment.

Eremogone kingii is extremely variable throughout its range with six infraspecific taxa recognized (under Arenaria kingii) by B. Maguire (1947, 1951). We have been unsuccessful in distinguishing more than two of those taxa. The others intergrade to such an extent that formal recognition is unwarranted. Most distinctive of these here-rejected taxa is var. uintahensis, said to have sepals (4.5–)5–6 mm, versus 3.6–4.5(–5) for the other taxa. Interestingly, the type specimen has sepals mostly 4.5 mm long. In the main portion of the range of var. uintahensis, the sepals are rounded to broadly obtuse, but they may also be acute. Furthermore, the sepals and pedicels are often glabrous, but the correlation of the above-mentioned characters varies over the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Caryophyllaceae > subfam. Alsinoideae > Eremogone Caryophyllaceae > subfam. Alsinoideae > Eremogone
Sibling taxa
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. franklinii, E. hookeri, E. macradenia, E. pumicola, E. stenomeres, E. ursina
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. hookeri, E. kingii, E. macradenia, E. pumicola, E. stenomeres, E. ursina
Subordinate taxa
E. kingii var. glabrescens, E. kingii var. kingii
E. franklinii var. franklinii, E. franklinii var. thompsonii
Key
1. Petals white, apex 2-fid
var. kingii
1. Petals white or rarely pink, apex entire or erose
var. glabrescens
1. Inflorescences tightly congested cymes; pedicels 0.1-1(-3) mm; sepals 8.5-12 mm; petals 0.8-0.9 times as long as sepals
var. franklinii
1. Inflorescences somewhat congested cymes; pedicels 1-4 mm; sepals 5-8 mm; petals 1-1.1 times as long as sepals
var. thompsonii
Synonyms Stellaria kingii, Arenaria kingii Arenaria franklinii
Name authority (S. Watson) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 140. (1973) (Douglas ex Hooker) R. L. Hartman & Rabeler: Sida 21: 240. (2004)
Source FNA vol. 5, p. 67. Treatment authors: Ronald L. Hartman, Richard K. Rabeler, Frederick H. Utech. FNA vol. 5, p. 66. Treatment authors: Ronald L. Hartman, Richard K. Rabeler, Frederick H. Utech.
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