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King's sandwort

Mount Dellanbaugh sandwort

Habit Plants tufted or sometimes in compact cushion, green, not glaucous, woody or not at base. Plants tufted to mat forming, green, not glaucous, with woody base.
Stems

erect, (1–)3–20(–25) cm, stipitate-glandular or glabrous proximally.

erect, (3–)10–23 cm, moderately to densely stipitate-glandular.

Leaves

basal leaves abundant, persistent;

cauline leaves in (1–)4+ pairs, reduced distally or not;

basal blades erect or closely ascending to somewhat spreading, green to gray-green, filiform to needlelike or narrowly subulate, 0.3–3(–4) cm × 0.3–1.2 mm, flexuous or rigid, herbaceous, apex apiculate or stiff and spinose, glabrous to stipitate-glandular, not glaucous.

basal leaves abundant, persistent;

cauline leaves in 5–7 pairs, reduced distally;

basal blades spreading to arcuate-spreading, needlelike, 0.8–2 cm × 0.4–0.8 mm, ± rigid, not fleshy, herbaceous, apex spinose, glabrous, not glaucous.

Inflorescences

(1–)3–13-flowered, ± open cymes.

(1–)3–6-flowered, open cymes.

Pedicels

2–15 mm, glabrous to densely stipitate-glandular.

6–25 mm, stipitate-glandular.

Flowers

sepals 1–3-veined, lateral veins less developed, ovate or lanceolate, (2.5–)2.8–5(–6) mm, not expanding in fruit, margins broad, apex obtuse to broadly acute or acuminate, glabrous or stipitate-glandular on herbaceous portion;

petals white or rarely pink, oblong to spatulate, (3–)4–7 mm, ca. 1.2–1.3 times as long as sepals, apex entire, erose, or 2-fid almost to base;

nectaries as abaxial, rounded lobe with transverse groove or elongate cup at base of filaments opposite sepals, 0.6 × 0.3 mm.

sepals 1–3-veined, lateral veins less developed, narrowly elliptic to ovate, 3.5–4 mm, 4.8–5.2 mm in fruit, margins usually broadly winged, scarious, apex broadly acute to obtuse (at least in fruit), glabrous or nearly so;

petals yellowish white, spatulate, 5.8–10 mm, 1.3–1.5 times as long as sepals, apex rounded;

nectaries as lateral and abaxial rounding of base of filaments opposite sepals, 0.2–0.3 mm.

Capsules

4.5–7 mm, glabrous.

7–10 mm, glabrous.

Seeds

black to brown, spheric or oblong to ovoid, 1.2–2.1 mm, low-tuberculate, sometimes papillate on abaxial ridge.

brownish black, suborbicular with hilar notch, 2–2.4 mm, tuberculate;

tubercles rounded, elongate.

Eremogone kingii

Eremogone aberrans

Phenology Flowering late spring–summer.
Habitat Oak and yellow pine forests
Elevation 1500-2800 m [4900-9200 ft]
Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

M. F. Baad (1969) considered Eremogone kingii to be monophyletic despite considerable morphological variation; J. C. Hickman (1971) thought otherwise, considering E. kingii to be a “genetic dumping ground for all the closely related taxa,” but did not propose any new taxonomic alignment.

Eremogone kingii is extremely variable throughout its range with six infraspecific taxa recognized (under Arenaria kingii) by B. Maguire (1947, 1951). We have been unsuccessful in distinguishing more than two of those taxa. The others intergrade to such an extent that formal recognition is unwarranted. Most distinctive of these here-rejected taxa is var. uintahensis, said to have sepals (4.5–)5–6 mm, versus 3.6–4.5(–5) for the other taxa. Interestingly, the type specimen has sepals mostly 4.5 mm long. In the main portion of the range of var. uintahensis, the sepals are rounded to broadly obtuse, but they may also be acute. Furthermore, the sepals and pedicels are often glabrous, but the correlation of the above-mentioned characters varies over the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eremogone aberrans is known only from northern Arizona and resembles a robust form of the more northerly occurring E. aculeata. In Arizona it is often confused with E. fendleri, which has sepals more or less glandular-pubescent whereas E. aberrans has sepals glabrous or with a few glandular hairs at their bases.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Caryophyllaceae > subfam. Alsinoideae > Eremogone Caryophyllaceae > subfam. Alsinoideae > Eremogone
Sibling taxa
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. franklinii, E. hookeri, E. macradenia, E. pumicola, E. stenomeres, E. ursina
E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. franklinii, E. hookeri, E. kingii, E. macradenia, E. pumicola, E. stenomeres, E. ursina
Subordinate taxa
E. kingii var. glabrescens, E. kingii var. kingii
Key
1. Petals white, apex 2-fid
var. kingii
1. Petals white or rarely pink, apex entire or erose
var. glabrescens
Synonyms Stellaria kingii, Arenaria kingii Arenaria aberrans
Name authority (S. Watson) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 140. (1973) (M. E. Jones) Ikonnikov: Novosti Sist. Vyssh. Rast. 10: 139. (1973)
Source FNA vol. 5, p. 67. Treatment authors: Ronald L. Hartman, Richard K. Rabeler, Frederick H. Utech. FNA vol. 5, p. 59. Treatment authors: Ronald L. Hartman, Richard K. Rabeler, Frederick H. Utech.
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