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Eremogone franklinii

Franklin's sandwort

King's sandwort

Habit Plants ± cespitose, bluish green, not glaucous, with woody base. Plants tufted or sometimes in compact cushion, green, not glaucous, woody or not at base.
Stems

erect, 3–10(–15) cm, glabrous.

erect, (1–)3–20(–25) cm, stipitate-glandular or glabrous proximally.

Leaves

basal leaves persistent;

cauline leaves in 6–10 pairs, closely overlapping, not reduced;

basal blades arcuate-spreading, needlelike, (0.6–)1–2 cm × ca. 1 mm, ± rigid, herbaceous, apex spinose, glabrous, not glaucous.

basal leaves abundant, persistent;

cauline leaves in (1–)4+ pairs, reduced distally or not;

basal blades erect or closely ascending to somewhat spreading, green to gray-green, filiform to needlelike or narrowly subulate, 0.3–3(–4) cm × 0.3–1.2 mm, flexuous or rigid, herbaceous, apex apiculate or stiff and spinose, glabrous to stipitate-glandular, not glaucous.

Inflorescences

3–45+-flowered, usually congested, subcapitate cymes.

(1–)3–13-flowered, ± open cymes.

Pedicels

ca. 0.1–3(–4) mm, glabrous.

2–15 mm, glabrous to densely stipitate-glandular.

Flowers

sepals 1–3-veined, linear-lanceolate, (5–)8.5–12 mm, not enlarging in fruit, margins narrow, apex acuminate, glabrous;

petals white, oblong-lanceolate to narrowly spatulate, 7–9 mm, 0.8–1.1 times as long as sepals, apex rounded to blunt;

nectaries not apparent.

sepals 1–3-veined, lateral veins less developed, ovate or lanceolate, (2.5–)2.8–5(–6) mm, not expanding in fruit, margins broad, apex obtuse to broadly acute or acuminate, glabrous or stipitate-glandular on herbaceous portion;

petals white or rarely pink, oblong to spatulate, (3–)4–7 mm, ca. 1.2–1.3 times as long as sepals, apex entire, erose, or 2-fid almost to base;

nectaries as abaxial, rounded lobe with transverse groove or elongate cup at base of filaments opposite sepals, 0.6 × 0.3 mm.

Capsules

2.3–3.3 mm, glabrous.

4.5–7 mm, glabrous.

Seeds

black, pyriform, 1.2–1.7 mm, tuberculate.

black to brown, spheric or oblong to ovoid, 1.2–2.1 mm, low-tuberculate, sometimes papillate on abaxial ridge.

Eremogone franklinii

Eremogone kingii

Distribution
from FNA
ID; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

M. F. Baad (1969) considered Eremogone kingii to be monophyletic despite considerable morphological variation; J. C. Hickman (1971) thought otherwise, considering E. kingii to be a “genetic dumping ground for all the closely related taxa,” but did not propose any new taxonomic alignment.

Eremogone kingii is extremely variable throughout its range with six infraspecific taxa recognized (under Arenaria kingii) by B. Maguire (1947, 1951). We have been unsuccessful in distinguishing more than two of those taxa. The others intergrade to such an extent that formal recognition is unwarranted. Most distinctive of these here-rejected taxa is var. uintahensis, said to have sepals (4.5–)5–6 mm, versus 3.6–4.5(–5) for the other taxa. Interestingly, the type specimen has sepals mostly 4.5 mm long. In the main portion of the range of var. uintahensis, the sepals are rounded to broadly obtuse, but they may also be acute. Furthermore, the sepals and pedicels are often glabrous, but the correlation of the above-mentioned characters varies over the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences tightly congested cymes; pedicels 0.1-1(-3) mm; sepals 8.5-12 mm; petals 0.8-0.9 times as long as sepals
var. franklinii
1. Inflorescences somewhat congested cymes; pedicels 1-4 mm; sepals 5-8 mm; petals 1-1.1 times as long as sepals
var. thompsonii
1. Petals white, apex 2-fid
var. kingii
1. Petals white or rarely pink, apex entire or erose
var. glabrescens
Source FNA vol. 5, p. 66. FNA vol. 5, p. 67.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Eremogone Caryophyllaceae > subfam. Alsinoideae > Eremogone
Sibling taxa
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. hookeri, E. kingii, E. macradenia, E. pumicola, E. stenomeres, E. ursina
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. franklinii, E. hookeri, E. macradenia, E. pumicola, E. stenomeres, E. ursina
Subordinate taxa
E. franklinii var. franklinii, E. franklinii var. thompsonii
E. kingii var. glabrescens, E. kingii var. kingii
Synonyms Arenaria franklinii Stellaria kingii, Arenaria kingii
Name authority (Douglas ex Hooker) R. L. Hartman & Rabeler: Sida 21: 240. (2004) (S. Watson) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 140. (1973)
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