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Eremogone franklinii

Franklin's sandwort

Habit Plants ± cespitose, bluish green, not glaucous, with woody base. Herbs, winter annual, annual, biennial, or perennial; taprooted and/or rhizomatous, rarely with tuberous thickenings (Pseudostellaria).
Stems

erect, 3–10(–15) cm, glabrous.

prostrate to ascending or erect, simple or branched.

Leaves

basal leaves persistent;

cauline leaves in 6–10 pairs, closely overlapping, not reduced;

basal blades arcuate-spreading, needlelike, (0.6–)1–2 cm × ca. 1 mm, ± rigid, herbaceous, apex spinose, glabrous, not glaucous.

opposite, connate proximally or not, often petiolate (basal leaves), not stipulate;

blade subulate or linear to spatulate, lanceolate, or broadly ovate, seldom succulent.

Inflorescences

3–45+-flowered, usually congested, subcapitate cymes.

terminal or axillary cymes, or flowers solitary;

bracts foliaceous or reduced, herbaceous to scarious (or rarely absent);

involucel bracteoles absent.

Pedicels

ca. 0.1–3(–4) mm, glabrous.

present or rarely flowers sessile.

Flowers

sepals 1–3-veined, linear-lanceolate, (5–)8.5–12 mm, not enlarging in fruit, margins narrow, apex acuminate, glabrous;

petals white, oblong-lanceolate to narrowly spatulate, 7–9 mm, 0.8–1.1 times as long as sepals, apex rounded to blunt;

nectaries not apparent.

bisexual or seldom unisexual, sometimes inconspicuous;

perianth and androecium hypogynous or perigynous, often slightly;

hypanthium cup-, dish-, or disc-shaped;

sepals (4–)5, distinct or seldom connate basally, sometimes hooded, not awned;

petals absent or (1–)4–5, usually white, sometimes translucent, yellowish white, pink, or brownish, seldom clawed, auricles absent, coronal appendages absent, blade apex entire or 2-fid, sometimes jagged or emarginate, rarely laciniate;

stamens absent or (1–)5(–10), in 1 or 2 whorls, arising from base of ovary, a nectariferous disc, or sometimes the hypanthium or hypanthium rim;

staminodes absent or 1–5(–8);

ovary 1- or rarely 3-locular (Wilhelmsia);

styles (2–)3–5(–6), distinct;

stigmas (2–)3–5(–6).

Fruits

capsules, or rarely utricles (Scleranthus), opening by (2–)3–6, occasionally 8 or 10 valves or (3 or) 6–10 teeth;

carpophore present or often absent.

Capsules

2.3–3.3 mm, glabrous.

Seeds

black, pyriform, 1.2–1.7 mm, tuberculate.

1–60+, yellowish or tan to dark red or often brown or black, usually reniform or triangular to circular and laterally compressed or ovoid to globose, rarely oblong and dorsiventrally compressed (Holosteum);

embryo usually peripheral and curved, rarely central and straight (Holosteum).

x

= 6–15, 17–19, 23.

Eremogone franklinii

Caryophyllaceae subfam. alsinoideae

Distribution
from FNA
ID; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
North-temperate regions; South America (Andean region); Europe (Mediterranean region); w Asia; c Asia (Himalayas, Mediterranean region); Africa (Mediterranean region)
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 30, species ca. 1040 (16 genera, 137 species in the flora).

Alsinoideae, often considered basal in the family and the least specialized, is in some ways the most heterogeneous of the subfamilies. Members of its largest tribe (Alsineae) share the following characteristics: stipules absent, sepals free or at most basally connate, and capsular fruits. Indehiscent fruits, relatively short hypanthia, and other floral reductions occur in varying combinations in the approximately 30 species placed in four other tribes. A broad molecular survey of Alsinoideae has revealed two major lineages and lack of support for the existing tribal circumscriptions (M. Nepokroeff et al. 2002). About three-fourths of the species are members of Arenaria, Cerastium, Minuartia, and Stellaria.

Attempts have been made to move Scleranthus (fruit a utricle surrounded by an enlarged hypanthium) from Alsinoideae to either Paronychioideae (J. Hutchinson 1973, as Illecebraceae) or Scleranthaceae (A. Takhtajan 1997). Recent molecular and morphological studies by R. D. Smissen et. al. (2002, 2003) supported its retention in the Alsinoideae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences tightly congested cymes; pedicels 0.1-1(-3) mm; sepals 8.5-12 mm; petals 0.8-0.9 times as long as sepals
var. franklinii
1. Inflorescences somewhat congested cymes; pedicels 1-4 mm; sepals 5-8 mm; petals 1-1.1 times as long as sepals
var. thompsonii
Source FNA vol. 5, p. 66. FNA vol. 5, p. 50. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Eremogone Caryophyllaceae
Sibling taxa
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. fendleri, E. ferrisiae, E. hookeri, E. kingii, E. macradenia, E. pumicola, E. stenomeres, E. ursina
Subordinate taxa
E. franklinii var. franklinii, E. franklinii var. thompsonii
Synonyms Arenaria franklinii
Name authority (Douglas ex Hooker) R. L. Hartman & Rabeler: Sida 21: 240. (2004) Fenzl: in S. L. Endlicher, Gen. Pl. 13: 963. (1840)
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