Eremogone fendleri |
Eremogone kingii |
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Fendler's sandwort |
King's sandwort |
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Habit | Plants ± cespitose, bluish green, not glaucous, with woody base. | Plants tufted or sometimes in compact cushion, green, not glaucous, woody or not at base. | ||||
Stems | erect, (2–)10–30(–40) cm, stipitate-glandular. |
erect, (1–)3–20(–25) cm, stipitate-glandular or glabrous proximally. |
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Leaves | basal leaves persistent; cauline leaves in (4–)5+ pairs, reduced or not; basal blades ascending or recurved, filiform, 1–10(–11) cm × 0.2–0.4 mm, flexuous, herbaceous, apex apiculate to spinose, glabrous to puberulent, not glaucous. |
basal leaves abundant, persistent; cauline leaves in (1–)4+ pairs, reduced distally or not; basal blades erect or closely ascending to somewhat spreading, green to gray-green, filiform to needlelike or narrowly subulate, 0.3–3(–4) cm × 0.3–1.2 mm, flexuous or rigid, herbaceous, apex apiculate or stiff and spinose, glabrous to stipitate-glandular, not glaucous. |
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Inflorescences | (1–)3–35-flowered, ± open cymes. |
(1–)3–13-flowered, ± open cymes. |
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Pedicels | 3–25 mm, stipitate-glandular. |
2–15 mm, glabrous to densely stipitate-glandular. |
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Flowers | sepals weakly to prominently 1–3-veined, linear-lanceolate, 4–7.5 mm, not enlarging in fruit, margins broad, apex acuminate, moderately to densely stipitate-glandular on herbaceous portion; petals white, oblong-elliptic to spatulate, 4–8 mm, 0.9–1.3 times as long as sepals, apex entire to somewhat erose; nectaries as lateral and abaxial rounding of base of filaments opposite sepals, 0.2 × 0.4 mm. |
sepals 1–3-veined, lateral veins less developed, ovate or lanceolate, (2.5–)2.8–5(–6) mm, not expanding in fruit, margins broad, apex obtuse to broadly acute or acuminate, glabrous or stipitate-glandular on herbaceous portion; petals white or rarely pink, oblong to spatulate, (3–)4–7 mm, ca. 1.2–1.3 times as long as sepals, apex entire, erose, or 2-fid almost to base; nectaries as abaxial, rounded lobe with transverse groove or elongate cup at base of filaments opposite sepals, 0.6 × 0.3 mm. |
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Capsules | 5–7 mm, glabrous. |
4.5–7 mm, glabrous. |
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Seeds | black, ovoid to pyriform with hilar notch, 1.5–1.9 mm, tuberculate; tubercles rounded, elongate to rounded-conic. |
black to brown, spheric or oblong to ovoid, 1.2–2.1 mm, low-tuberculate, sometimes papillate on abaxial ridge. |
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2n | = 44. |
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Eremogone fendleri |
Eremogone kingii |
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Phenology | Flowering spring–late summer. | |||||
Habitat | Sagebrush plains, pine forests, and mountain slopes to alpine zones | |||||
Elevation | 1200-4300 m (3900-14100 ft) | |||||
Distribution |
AZ; CO; NM; TX; UT; WY
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CA; ID; NV; OR; UT
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Discussion | We agree with M. F. Baad (1969) in not formally recognizing varieties within Eremogone fendleri. B. Maguire (1947, 1951) recognized five varieties, defined chiefly on leaf and sepal characteristics. While some specimens can be “matched” to varieties, many appear intermediate between them, forming a continuum of variation. B. Maguire (1947) noted that Eremogone fendleri is “probably to be found in the states of Sonora and Chihuahua, Mexico”; we have not seen any collections from that area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). M. F. Baad (1969) considered Eremogone kingii to be monophyletic despite considerable morphological variation; J. C. Hickman (1971) thought otherwise, considering E. kingii to be a “genetic dumping ground for all the closely related taxa,” but did not propose any new taxonomic alignment. Eremogone kingii is extremely variable throughout its range with six infraspecific taxa recognized (under Arenaria kingii) by B. Maguire (1947, 1951). We have been unsuccessful in distinguishing more than two of those taxa. The others intergrade to such an extent that formal recognition is unwarranted. Most distinctive of these here-rejected taxa is var. uintahensis, said to have sepals (4.5–)5–6 mm, versus 3.6–4.5(–5) for the other taxa. Interestingly, the type specimen has sepals mostly 4.5 mm long. In the main portion of the range of var. uintahensis, the sepals are rounded to broadly obtuse, but they may also be acute. Furthermore, the sepals and pedicels are often glabrous, but the correlation of the above-mentioned characters varies over the range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 64. | FNA vol. 5, p. 67. | ||||
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Eremogone | Caryophyllaceae > subfam. Alsinoideae > Eremogone | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Arenaria fendleri, Arenaria fendleri subsp. brevifolia, Arenaria fendleri var. brevifolia, Arenaria fendleri var. diffusa, Arenaria fendleri var. porteri, Arenaria fendleri var. tweedyi, Arenaria tweedyi | Stellaria kingii, Arenaria kingii | ||||
Name authority | (A. Gray) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 139. (1973) | (S. Watson) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 140. (1973) | ||||
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