Eremogone fendleri |
Eremogone ferrisiae |
|
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Fendler's sandwort |
Ferris' sandwort |
|
Habit | Plants ± cespitose, bluish green, not glaucous, with woody base. | Plants tufted, green, not glaucous, with woody base. |
Stems | erect, (2–)10–30(–40) cm, stipitate-glandular. |
erect, (10–)20–40(–100) cm, glabrous to stipitate-glandular. |
Leaves | basal leaves persistent; cauline leaves in (4–)5+ pairs, reduced or not; basal blades ascending or recurved, filiform, 1–10(–11) cm × 0.2–0.4 mm, flexuous, herbaceous, apex apiculate to spinose, glabrous to puberulent, not glaucous. |
basal leaves sparse or absent; cauline leaves usually in 5–7 pairs, not significantly reduced; basal blades ascending, needlelike or narrowly linear, 2–6(–7) cm × 0.5–1 mm, ± rigid, herbaceous to subsucculent, apex blunt to spinose, usually glabrous, not glaucous. |
Inflorescences | (1–)3–35-flowered, ± open cymes. |
10–30(–80)-flowered, diffuse cymes; branches spreading. |
Pedicels | 3–25 mm, stipitate-glandular. |
15–55 mm, glabrous or stipitate-glandular. |
Flowers | sepals weakly to prominently 1–3-veined, linear-lanceolate, 4–7.5 mm, not enlarging in fruit, margins broad, apex acuminate, moderately to densely stipitate-glandular on herbaceous portion; petals white, oblong-elliptic to spatulate, 4–8 mm, 0.9–1.3 times as long as sepals, apex entire to somewhat erose; nectaries as lateral and abaxial rounding of base of filaments opposite sepals, 0.2 × 0.4 mm. |
sepals 1–3-veined, ovate to lanceolate or elliptic, 3–4.3 mm, to 5.5 mm in fruit, margins narrow to broad, apex acute to acuminate, glabrous to sparsely stipitate-glandular; petals white or yellowish, oblanceolate to spatulate, 6–9 mm, 1–1.5 times as long as sepals, apex entire or erose; nectaries as lateral and abaxial rounding of base of filaments opposite sepals, 0.3–0.4 mm. |
Capsules | 5–7 mm, glabrous. |
6–7 mm, glabrous. |
Seeds | black, ovoid to pyriform with hilar notch, 1.5–1.9 mm, tuberculate; tubercles rounded, elongate to rounded-conic. |
reddish brown to blackish, suborbicular to pyriform or ovoid, 1.3–3.2 mm, tuberculate; tubercles low, rounded to conic. |
2n | = 44. |
|
Eremogone fendleri |
Eremogone ferrisiae |
|
Phenology | Flowering spring–late summer. | Flowering spring–summer. |
Habitat | Sagebrush plains, pine forests, and mountain slopes to alpine zones | Pine and oak woodlands, granitic alluvium on foothills and mountain slopes |
Elevation | 1200-4300 m (3900-14100 ft) | 1400-2900 m (4600-9500 ft) |
Distribution |
AZ; CO; NM; TX; UT; WY
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AZ; CA; NV; UT
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Discussion | We agree with M. F. Baad (1969) in not formally recognizing varieties within Eremogone fendleri. B. Maguire (1947, 1951) recognized five varieties, defined chiefly on leaf and sepal characteristics. While some specimens can be “matched” to varieties, many appear intermediate between them, forming a continuum of variation. B. Maguire (1947) noted that Eremogone fendleri is “probably to be found in the states of Sonora and Chihuahua, Mexico”; we have not seen any collections from that area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
We now believe that Eremogone macradenia (in the sense of R. L. Hartman 1993) should be split into two species, with E. macradenia var. ferrisiae (Abrams) R. L. Hartman & Rabeler being elevated to species rank (Hartman and R. K. Rabeler 2004), as here. This became particularly obvious when comparing nectary morphology of E. macradenia (rectangular, two-lobed or truncate, 0.7–1.5 mm or narrowly longitudinally rectangular, truncate, densely minutely pubescent with erect to spreading hairs, 0.7–0.8 mm) with that of E. ferrisiae (rounded, 0.3–0.4 mm). Furthermore, the nectary types correlate well with sepal size and inflorescence type, as indicated in the key. This disposition agrees with the conclusions of M. F. Baad (1969). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 64. | FNA vol. 5, p. 65. |
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Eremogone | Caryophyllaceae > subfam. Alsinoideae > Eremogone |
Sibling taxa | ||
Synonyms | Arenaria fendleri, Arenaria fendleri subsp. brevifolia, Arenaria fendleri var. brevifolia, Arenaria fendleri var. diffusa, Arenaria fendleri var. porteri, Arenaria fendleri var. tweedyi, Arenaria tweedyi | Arenaria macradenia subsp. ferrisiae, E. macradenia var. ferrisiae |
Name authority | (A. Gray) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 139. (1973) | (Abrams) R. L. Hartman & Rabeler: Sida 21: 754. (2004) |
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