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Fendler's sandwort

Eastwood's sandwort

Habit Plants ± cespitose, bluish green, not glaucous, with woody base. Plants densely matted, green, not glaucous, with woody base.
Stems

erect, (2–)10–30(–40) cm, stipitate-glandular.

erect, (8–)10–25 cm, glabrous or stipitate-glandular.

Leaves

basal leaves persistent;

cauline leaves in (4–)5+ pairs, reduced or not;

basal blades ascending or recurved, filiform, 1–10(–11) cm × 0.2–0.4 mm, flexuous, herbaceous, apex apiculate to spinose, glabrous to puberulent, not glaucous.

basal leaves persistent;

cauline leaves usually in 2–4 pairs, reduced distally;

basal blades spreading to recurved, needlelike, 1–3(–3.5) cm × 0.5–0.7 mm, flexuous to rigid, herbaceous, apex spinose, glabrous to puberulent, not glaucous.

Inflorescences

(1–)3–35-flowered, ± open cymes.

(1–)3–17-flowered, ± open cymes.

Pedicels

3–25 mm, stipitate-glandular.

3–30 mm, glabrous or stipitate-glandular.

Flowers

sepals weakly to prominently 1–3-veined, linear-lanceolate, 4–7.5 mm, not enlarging in fruit, margins broad, apex acuminate, moderately to densely stipitate-glandular on herbaceous portion;

petals white, oblong-elliptic to spatulate, 4–8 mm, 0.9–1.3 times as long as sepals, apex entire to somewhat erose;

nectaries as lateral and abaxial rounding of base of filaments opposite sepals, 0.2 × 0.4 mm.

sepals green or purplish, 1–3-veined, lanceolate to ovate-lanceolate, (3.5–)4–6.5 mm, not enlarging in fruit, margins broad, apex narrowly acute to acuminate, glabrous or stipitate-glandular;

petals yellowish white or sometimes brownish to reddish pink, broadly oblong-elliptic to oblanceolate, 4–6.5 mm, 0.9–1.1 times as long as sepals, apex rounded;

nectaries narrowly longitudinally rectangular, apically cleft or emarginate, adjacent to filaments opposite sepals, 1–2 mm.

Capsules

5–7 mm, glabrous.

4–6 mm, glabrous.

Seeds

black, ovoid to pyriform with hilar notch, 1.5–1.9 mm, tuberculate;

tubercles rounded, elongate to rounded-conic.

brown, ovoid to suborbicular with hilar notch, 1.2–1.7 mm, papillate, subechinate;

tubercles conical.

2n

= 44.

Eremogone fendleri

Eremogone eastwoodiae

Phenology Flowering spring–late summer.
Habitat Sagebrush plains, pine forests, and mountain slopes to alpine zones
Elevation 1200-4300 m (3900-14100 ft)
Distribution
from FNA
AZ; CO; NM; TX; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

We agree with M. F. Baad (1969) in not formally recognizing varieties within Eremogone fendleri. B. Maguire (1947, 1951) recognized five varieties, defined chiefly on leaf and sepal characteristics. While some specimens can be “matched” to varieties, many appear intermediate between them, forming a continuum of variation.

B. Maguire (1947) noted that Eremogone fendleri is “probably to be found in the states of Sonora and Chihuahua, Mexico”; we have not seen any collections from that area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The Hopi Indians may use Eremogone eastwoodiae as an emetic (B. Maguire 1960).

The nectaries in Eremogone eastwoodiae are different from those of most other species of the genus in North America since they are a separate bilobed structure adjacent to, but not a direct enlargement of, the filament bases opposite the sepals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems and pedicels glabrous
var. eastwoodiae
1. Stems and pedicels stipitate-glandular
var. adenophora
Source FNA vol. 5, p. 64. FNA vol. 5, p. 63.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Eremogone Caryophyllaceae > subfam. Alsinoideae > Eremogone
Sibling taxa
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. eastwoodiae, E. ferrisiae, E. franklinii, E. hookeri, E. kingii, E. macradenia, E. pumicola, E. stenomeres, E. ursina
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. fendleri, E. ferrisiae, E. franklinii, E. hookeri, E. kingii, E. macradenia, E. pumicola, E. stenomeres, E. ursina
Subordinate taxa
E. eastwoodiae var. adenophora, E. eastwoodiae var. eastwoodiae
Synonyms Arenaria fendleri, Arenaria fendleri subsp. brevifolia, Arenaria fendleri var. brevifolia, Arenaria fendleri var. diffusa, Arenaria fendleri var. porteri, Arenaria fendleri var. tweedyi, Arenaria tweedyi Arenaria eastwoodiae, Arenaria fendleri var. eastwoodiae
Name authority (A. Gray) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 139. (1973) (Rydberg) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 139. (1973)
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