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Eastwood's sandwort

Habit Plants densely matted, green, not glaucous, with woody base. Herbs, winter annual, annual, biennial, or perennial; taprooted and/or rhizomatous, rarely with tuberous thickenings (Pseudostellaria).
Stems

erect, (8–)10–25 cm, glabrous or stipitate-glandular.

prostrate to ascending or erect, simple or branched.

Leaves

basal leaves persistent;

cauline leaves usually in 2–4 pairs, reduced distally;

basal blades spreading to recurved, needlelike, 1–3(–3.5) cm × 0.5–0.7 mm, flexuous to rigid, herbaceous, apex spinose, glabrous to puberulent, not glaucous.

opposite, connate proximally or not, often petiolate (basal leaves), not stipulate;

blade subulate or linear to spatulate, lanceolate, or broadly ovate, seldom succulent.

Inflorescences

(1–)3–17-flowered, ± open cymes.

terminal or axillary cymes, or flowers solitary;

bracts foliaceous or reduced, herbaceous to scarious (or rarely absent);

involucel bracteoles absent.

Pedicels

3–30 mm, glabrous or stipitate-glandular.

present or rarely flowers sessile.

Flowers

sepals green or purplish, 1–3-veined, lanceolate to ovate-lanceolate, (3.5–)4–6.5 mm, not enlarging in fruit, margins broad, apex narrowly acute to acuminate, glabrous or stipitate-glandular;

petals yellowish white or sometimes brownish to reddish pink, broadly oblong-elliptic to oblanceolate, 4–6.5 mm, 0.9–1.1 times as long as sepals, apex rounded;

nectaries narrowly longitudinally rectangular, apically cleft or emarginate, adjacent to filaments opposite sepals, 1–2 mm.

bisexual or seldom unisexual, sometimes inconspicuous;

perianth and androecium hypogynous or perigynous, often slightly;

hypanthium cup-, dish-, or disc-shaped;

sepals (4–)5, distinct or seldom connate basally, sometimes hooded, not awned;

petals absent or (1–)4–5, usually white, sometimes translucent, yellowish white, pink, or brownish, seldom clawed, auricles absent, coronal appendages absent, blade apex entire or 2-fid, sometimes jagged or emarginate, rarely laciniate;

stamens absent or (1–)5(–10), in 1 or 2 whorls, arising from base of ovary, a nectariferous disc, or sometimes the hypanthium or hypanthium rim;

staminodes absent or 1–5(–8);

ovary 1- or rarely 3-locular (Wilhelmsia);

styles (2–)3–5(–6), distinct;

stigmas (2–)3–5(–6).

Fruits

capsules, or rarely utricles (Scleranthus), opening by (2–)3–6, occasionally 8 or 10 valves or (3 or) 6–10 teeth;

carpophore present or often absent.

Capsules

4–6 mm, glabrous.

Seeds

brown, ovoid to suborbicular with hilar notch, 1.2–1.7 mm, papillate, subechinate;

tubercles conical.

1–60+, yellowish or tan to dark red or often brown or black, usually reniform or triangular to circular and laterally compressed or ovoid to globose, rarely oblong and dorsiventrally compressed (Holosteum);

embryo usually peripheral and curved, rarely central and straight (Holosteum).

x

= 6–15, 17–19, 23.

Eremogone eastwoodiae

Caryophyllaceae subfam. alsinoideae

Distribution
from FNA
AZ; CO; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
North-temperate regions; South America (Andean region); Europe (Mediterranean region); w Asia; c Asia (Himalayas, Mediterranean region); Africa (Mediterranean region)
Discussion

Varieties 2 (2 in the flora).

The Hopi Indians may use Eremogone eastwoodiae as an emetic (B. Maguire 1960).

The nectaries in Eremogone eastwoodiae are different from those of most other species of the genus in North America since they are a separate bilobed structure adjacent to, but not a direct enlargement of, the filament bases opposite the sepals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 30, species ca. 1040 (16 genera, 137 species in the flora).

Alsinoideae, often considered basal in the family and the least specialized, is in some ways the most heterogeneous of the subfamilies. Members of its largest tribe (Alsineae) share the following characteristics: stipules absent, sepals free or at most basally connate, and capsular fruits. Indehiscent fruits, relatively short hypanthia, and other floral reductions occur in varying combinations in the approximately 30 species placed in four other tribes. A broad molecular survey of Alsinoideae has revealed two major lineages and lack of support for the existing tribal circumscriptions (M. Nepokroeff et al. 2002). About three-fourths of the species are members of Arenaria, Cerastium, Minuartia, and Stellaria.

Attempts have been made to move Scleranthus (fruit a utricle surrounded by an enlarged hypanthium) from Alsinoideae to either Paronychioideae (J. Hutchinson 1973, as Illecebraceae) or Scleranthaceae (A. Takhtajan 1997). Recent molecular and morphological studies by R. D. Smissen et. al. (2002, 2003) supported its retention in the Alsinoideae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems and pedicels glabrous
var. eastwoodiae
1. Stems and pedicels stipitate-glandular
var. adenophora
Source FNA vol. 5, p. 63. FNA vol. 5, p. 50. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Eremogone Caryophyllaceae
Sibling taxa
E. aberrans, E. aculeata, E. capillaris, E. congesta, E. fendleri, E. ferrisiae, E. franklinii, E. hookeri, E. kingii, E. macradenia, E. pumicola, E. stenomeres, E. ursina
Subordinate taxa
E. eastwoodiae var. adenophora, E. eastwoodiae var. eastwoodiae
Synonyms Arenaria eastwoodiae, Arenaria fendleri var. eastwoodiae
Name authority (Rydberg) Ikonnikov: Novosti Syst. Vyssh. Rast. 10: 139. (1973) Fenzl: in S. L. Endlicher, Gen. Pl. 13: 963. (1840)
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