1. Plants annual, tufted or mat-forming, without innovations [for opposite lead, see p. 67]. | → 2 |
2. Palea keels prominently ciliate, the cilia 0.2-0.8 mm long. | → 3 |
3. Spikelets 1-3.6 mm long, 0.9-2 mm wide, with 4-12 florets; lemmas 0.7-1.3 mm long. | → 4 |
4. Anthers 2; pedicels 0.1-1 mm long, mostly shorter than the spikelets, straight | E. ciliaris |
4. Anthers 3; pedicels 1-4(7) mm long, as long as or longer than the spikelets, mostly curved | E. amabilis |
3. Spikelets 5-20 mm long, 1.4-4 mm wide, with 10-42 florets; lemmas 1.3-2.8 mm long. | → 5 |
5. Lemmas and culms without glands; anthers 0.1-0.2 mm long, purplish | E. cumingii |
5. Lemmas with 1-3 crateriform glands on the keels, similar glands also often present below the cauline nodes; anthers 0.2-0.5 mm long, yellow | E. cilianensis |
2. Palea keels smooth or scabrous, the scabridities less than 0.2 mm long. | → 6 |
6. Plants mat-forming; panicles 1-3.5 cm long; erect portion of culms (2)5-20 cm, the basal portion prostrate and rooting at the nodes. | → 7 |
7. Spikelets bisexual; anthers 2, 0.2-0.3 mm long | E. hypnoides |
7. Spikelets and plants unisexual; anthers 3, 1.4-2.2 mm long | E. reptans |
6. Plants usually not forming mats; panicles 3-55 cm long; culms (2)6-130 cm tall, not prostrate or rooting at the lower nodes. | → 8 |
8. Ligules membranous, neither ciliolate nor ciliate | E. japonica |
8. Ligules membranous and ciliolate to ciliate, the cilia often longer than the basal membrane. | → 9 |
9. Caryopses with a shallow or deep ventral groove, ovoid to rectangular-prismatic or dorsally compressed, if dorsally compressed, the surface striate or smooth. | → 10 |
10. Bases of the caryopses greenish; caryopses dorsally compressed, the distal 2/3 translucent, the surface smooth; leaf sheaths with oblong glands; in the Flora region, known from a single collection at Canton, Maryland | E. cylindriflora |
10. Bases of the caryopses reddish-brown or brownish; caryopses laterally compressed or rectangular-prismatic to ovoid, the distal 2/3 opaque, the surface striate; sheaths without oblong glands, sometimes with glandular pits; plants found at many locations in the Flora region. | → 11 |
11. Spikelets 4-11 mm long, with 5-15 florets; pedicels somewhat divergent to almost appressed | E. mexicana |
11. Spikelets 1.4-5 mm long, with 2-7 florets; pedicels divergent. | → 12 |
12. Panicles 4-20 cm long, less than 1/2 the height of the plant; pedicels 1.5-5 mm long; glandular pits often present below the cauline nodes, on the rachises, and on the panicle branches | E. frankii |
12. Panicles 10-45(55) cm long, 2/3 or more the height of the plant; pedicels 4-25 mm long; plants without glandular pits | E. capillaris |
9. Caryopses without a ventral groove, usually globose, rarely flattened, pyriform, obovoid, ellipsoid, or rectangular-prismatic, the surface smooth to faintly striate. | → 13 |
13. Plants with glandular pits or bands somewhere, the location(s) various, including any or all of the following: below the cauline nodes, on the sheaths, blades, rachises, panicle branches, or pedicels, or on the keels of the lemmas and paleas. | → 14 |
14. Panicles 0.5-2 cm wide, contracted; primary panicle branches usually appressed, occasionally diverging up to 30° from the rachises; spikelets light yellowish, occasionally with reddish-purple markings | E. lutescens |
14. Panicles 2-18 cm wide, open to somewhat contracted; primary panicle branches diverging 20-110° from the rachises; spikelets plumbeous, greenish, or reddish-purple. | → 15 |
15. Spikelets 1.7-5.6 mm long, with 3-6 florets | E. frankii |
15. Spikelets (2)3.5-20 mm long, with (3)5-40 florets. | → 16 |
16. Spikelets 0.6-1.4 mm wide; pedicels 1-10 mm long, lax, appressed or divergent | E. pilosa |
16. Spikelets 1.1-4 mm wide; pedicels 0.2-4 mm long, stiff, straight, usually divergent. | → 17 |
17. Lemmas 2-2.8 mm long, with 1-3 crateriform glands along the keels; spikelets 6-20 mm long, 2-4 mm wide, with 10-40 florets; disarticulation below the florets, the rachillas persistent; anthers yellow | E. cilianensis |
17. Lemmas 1.4-1.8 mm long, rarely with 1 or 2 crateriform glands along the keels; spikelets 4-7(11) mm long, 1.1-2.2 mm wide, with 7-12(20) florets; disarticulation below the lemmas, both the paleas and rachillas usually persistent; anthers reddish-brown. | → 18 |
18. Panicles with glandular regions below the nodes, the glandular tissue forming a ring or band, often shiny or yellowish; anthers 3; blade margins without crateriform glands; pedicels without glandular bands | E. barrelieri |
18. Panicles sometimes with areas, but rarely rings, of glandular spots or crateriform pits below the nodes, the glands usually dull greenish-gray to stramineous; anthers 2; blade margins sometimes with crateriform glands; pedicels usually with glandular bands | E. minor |
13. Plants without glandular pits or bands. | → 19 |
19. Spikelets (1.6)2-4 mm wide; florets disarticulating intact from the persistent rachillas | E. unioloides |
19. Spikelets 0.6-2.5 mm wide; lemmas disarticulating separately from the paleas, sometimes both the paleas and the rachillas persistent. | → 20 |
20. Spikelets with 3-6 florets; plants of the central and northeastern United States and southern Ontario, Canada | E. frankii |
20. Spikelets with (3)5-42 florets; plants from throughout the contiguous United States and southern Ontario, Canada. | → 21 |
21. Lemmas 1.6-3 mm long; caryopses 0.7-1.3 mm long, obovoid, smooth, light brown to white; plants cultivated, occasionally escaping | E. tef |
21. Lemmas 1-2.2 mm long; caryopses 0.3-1.1 mm long, subglobose, pyriform, or obovoid to prism-shaped, smooth or faintly striate, brownish; plants native species or established introductions, variously distributed. | → 22 |
22. Lemmas with conspicuous, often greenish lateral veins; caryopses 0.3-0.6 mm long, ovoid, subglobose to obovoid. | → 23 |
23. Spikelets 5-12(18) mm long, with 12-42 florets; primary branches 6-10 per culm; lemmas 1.3-2 mm long; anthers 3 | E. cumingii |
23. Spikelets 2-4.6 mm long, with 5-15 florets; primary branches (12)15-20 per culm; lemmas 1-1.3 mm long; anthers 2 | E. gangetica |
22. Lemmas with inconspicuous to moderately conspicuous lateral veins, the veins usually not greenish; caryopses 0.5-1.1 mm long, pyriform or obovoid to prism-shaped. | → 24 |
24. Lower glumes 0.5-1.5 mm long, at least 1/2 as long as the lowest lemmas; spikelets 1.2-2.5 mm wide; panicle branches solitary or paired at the lowest 2 nodes; lemmas with moderately conspicuous lateral veins | E. pectinacea |
24. Lower glumes 0.3-0.6(0.8) mm long, usually less than 1/2 as long as the lowest lemmas; spikelets 0.6-1.4 mm wide; panicle branches usually whorled at the lowest 2 nodes; lemmas with inconspicuous lateral veins | E. pilosa |
1. Plants perennial, sometimes rhizomatous, forming innovations at the basal nodes [for opposite lead, see p. 65]. | → 25 |
25. Paleas with a broad lower portion forming a wing or tooth on each side, these often projecting beyond the lemmas. | → 26 |
26. Spikelets 5.5-16 mm long, 2.7-9 mm wide; lemmas 3-5 mm long, the keels without crateriform glands; pedicels with a narrow band or abscission line just below the apices; anthers 1.4-2.8 mm long | E. superba |
26. Spikelets 2-5 mm long, 2-3.5 mm wide; lemmas 1.8-2.3 mm long, the keels with a few crateriform glands; pedicels without a narrow band or abscission line just below the apices; anthers 0.5-0.9 mm long | E. echinochloidea |
25. Paleas without a broad lower portion forming a wing or tooth, the bases never projecting beyond the lemmas. | → 27 |
27. Plants rhizomatous; disarticulation always below the florets, the paleas falling with the lemmas and caryopses. | → 28 |
28. Plants with long, scaly rhizomes, 4-8 mm thick; spikelets 8-14 mm long; lemmas 3.8-4.5 mm long, 3-5-veined, the apices acute to obtuse, usually erose; caryopses 1.6-2 mm long | E. obtusiflora |
28. Plants with short, knotty rhizomes less than 4 mm thick, often stout but never elongated; spikelets 2.5-7.6 mm long; lemmas 1-2.5 mm long, 3-veined, the apices acute, usually entire; caryopses 0.5-0.8 mm long. | → 29 |
29. Sheaths, blades, and culms not viscid or glandular; caryopses strongly flattened, the ventral surface with 2 prominent ridges separated by a groove; anthers 0.3-0.5 mm long; lemmas leathery | E. spectabilis |
29. Sheaths, blades, and/or culms often viscid, sometimes glandular; caryopses terete, the ventral surfaces without 2 ridges separated by a groove; anthers 0.2-0.4 mm long; lemmas membranous. | → 30 |
30. Pedicels 0.2-1.2 mm long, appressed; lemmas 1.5-2.2 mm long; caryopses 0.6-0.8 mm long | E. curtipedicellata |
30. Pedicels (1)1.5-12 mm long, divergent or appressed; lemmas 1.1-1.4 mm long; caryopses 0.5-0.6 mm long | E. silveana |
27. Plants not rhizomatous; disarticulation often below the lemmas, the paleas persistent, sometimes below the florets and the the paleas falling with the lemmas and caryopses. | → 31 |
31. Panicles 0.3-0.6 cm wide, spicate, dense; spikelets with 2-3 florets | E. spicata |
31. Panicles 1-45 cm wide, ovate to obovate or elliptic, open to somewhat condensed and glomerate; spikelets with 1-45 florets. | → 32 |
32. Caryopses with shallowly to deeply grooved adaxial surfaces, rectangular-prismatic to ellipsoid, ovoid, or obovoid in overall shape [for opposite lead, see p. 69]. | → 33 |
33. Caryopses strongly dorsally compressed, translucent, mostly light brown, bases sometimes greenish. | → 34 |
34. Lemmas 1.8-3 mm long; panicles 16-35(40) cm long, (4)8-24 cm wide; blades 12-50(65) cm long; caryopses 1-1.7 mm long; ligules 0.6-1.3 mm long | E. curvula |
34. Lemmas 1.4-1.7 long; panicles 6-18 cm long, 2-8 cm wide; blades 2-12 cm long; caryopses 0.4-0.8 mm long; ligules 0.3-0.5 mm long. | → 35 |
35. Plants without woolly hairs at the base; glumes unequal; lateral lemma veins not green, inconspicuous throughout; spikelets 0.8-1.2 mm; naturalized in the southwestern United States | E. lehmanniana |
35. Plants with conspicuous, woolly hairs at the base; glumes subequal; lateral lemma veins green, conspicuous basally, obscure near the lemma apices; spikelets 1.3-2 mm wide; in the Flora region, known only from waste areas near a woolen mill in South Carolina | E. setifolia |
33. Caryopses laterally compressed, terete, or slightly dorsally compressed, usually opaque, usually reddish-brown. | → 36 |
36. Lateral veins of the lemmas conspicuous, often greenish, the lemmas strongly keeled. | → 37 |
37. Panicles 2-8 cm wide, contracted to somewhat open, narrowly oblong to narrowly lanceolate; primary branches appressed or diverging up to 30° from the rachises; lemmas with punctate glands along the keels; pedicels 1-7 mm long, appressed; plants native to Africa, in the Flora region, known only from waste areas near sheep and cattle lots in South Carolina and Alabama | E. plana |
37. Panicles 4-30 cm wide, open, ovate to oblong; primary branches diverging 10-90° from the rachises; lemmas without punctate glands on the keels; pedicels 0.4-22 mm long, usually diverging, occasionally appressed; plants native to the southern United States. | → 38 |
38. Pedicels with a glandular band; culms with a glandular band below the nodes; anthers 0.3-0.5 mm long; restricted to southern Texas | E. swallenii |
38. Pedicels and culms without glandular bands; anthers 0.6-1.6 mm long; often found outside southern Texas. | → 39 |
39. Glumes 1.8-4 mm long, the upper glumes generally equaling or exceeding the lower lemmas; spikelets 1.5-3.6 mm wide, greenish-yellow with a reddish-purple tinge; lemmas 2.2-3.5 mm long; caryopses 0.8-1.3 mm long | E. trichodes |
39. Glumes 1.1-2.2 mm long, the upper glumes exceeded by the lower lemmas; spikelets 1-2 mm wide, plumbeous; lemmas 2-2.6 mm long; caryopses 0.6-0.8 mm long | E. palmeri |
36. Lateral veins of the lemmas inconspicuous, the lemmas sometimes only weakly keeled. | → 40 |
40. Lemmas 1.2-1.8 mm long; culms 30-70 cm tall. | → 41 |
41. Culms with a glandular ring below the nodes; bases of primary panicle branches with a glandular band; panicles 2-7 cm wide; pedicels glandular; known, in the Flora region, only from a few collections at Canton, Maryland | E. trichophora |
41. Culms without a glandular ring below the nodes; bases of primary panicle branches without a glandular band; pedicels not glandular at the base; panicles 5-27 cm wide; plants known from many parts of the southern United States. | → 42 |
42. Spikelets 1.1-1.6 mm wide, uniformly plumbeous; sheaths sometimes densely pilose dorsally and on the collars; distal margins of the lemmas not hyaline | E. polytricha |
42. Spikelets 0.5-1(1.3) mm wide, plumbeous to reddish-purple; sheaths usually glabrous dorsally and on the collars; distal margins of the lemmas hyaline | E. lugens |
40. Lemmas 1.6-3 mm long; culms (30)40-110(120) cm tall. | → 43 |
43. Spikelets greenish with a purplish tinge, with 2-6 florets; blades 25-60 cm long, 3-11 mm wide, flat to loosely involute; sheaths densely hirsute with papillose-based hairs on the collar, back, and base | E. hirsuta |
43. Spikelets olivaceous to plumbeous, with (3)5-12 florets; blades (4)10-35 cm long, 1-3.8 mm wide, involute or flat; sheaths never with papillose-based hairs, sometimes villous over the back. | → 44 |
44. Lemmas 1.6-2.2 mm long; anthers 0.5-0.8 mm long, purplish | E. intermedia |
44. Lemmas 2-3 mm long; anthers 0.6-1.7 mm long, purplish to yellowish. | → 45 |
45. Caryopses 0.8-1.6 mm long; lemmas 2.4-3 mm long | E. erosa |
45. Caryopses 0.6-0.8 mm long; lemmas 2-2.6 mm long | E. palmeri |
32. Caryopses not grooved on the adaxial surfaces, ellipsoid, subellipsoid, ovoid, obovoid, globose, to pyriform in overall shape [for opposite lead, see p. 68]. | → 46 |
| → 47 |
47. Panicles 15-45 cm wide, open, diffuse, broadly ovate to obovate; primary branches lax; pedicels 0.5-35(50) mm long, the lower pedicels longer or shorter than the spikelets. | → 48 |
48. Spikelets with appressed pedicels; only the terminal pedicels of each branch longer than the spikelets; disarticulation usually in the rachilla beneath the florets | E. refracta |
48. Spikelets with divergent pedicels; all pedicels usually longer than the spikelets; disarticulation below the lemmas, the paleas persistent | E. elliottii |
47. Panicles (1)2-17 cm wide, contracted to open, narrowly ovate to oblong; primary branches stiff; pedicels absent or 0.3-6 mm long, always shorter than the spikelets. | → 49 |
49. Spikelets 2.4-5 mm wide; glumes 1.7-4 mm long; lemmas 2-6 mm long, the apices usually acuminate or attenuate | E. secundiflora |
49. Spikelets 1-2.4 mm wide; glumes 1-2.2 mm long; lemmas 1.1-2.5 mm long, the apices usually acute, occasionally acuminate. | → 50 |
50. Spikelets 0.7-1.4 mm wide; anthers 0.2-0.3 mm long; caryopses flattened ventrally | E. prolifera |
50. Spikelets 1.3-2.4 mm wide; anthers (0.2)0.3-0.7 mm long; caryopses rounded, not flattened ventrally. | → 51 |
51. Terminal panicles 1-3.5 cm wide, contracted, condensed into glomerate lobes; primary branches 0.8-3 cm long | E. elongata |
51. Terminal panicles (1)2-17 cm wide, open to contracted; primary branches 1-15 cm long. | → 52 |
52. Plants without axillary panicles; terminal panicles 15-45 cm long; blades (8)12-40 cm long, 2-5 mm wide, flat to involute; caryopses 0.6-0.8 mm long, striate, obovoid to ellipsoid | E. bahiensis |
52. Plants usually with axillary panicles, these contracted and partially to completely enclosed by the subtending sheaths; terminal panicles 5-15 cm long; blades 4-8(18) cm long, 1-2 mm wide, usually involute; caryopses 0.5-0.6 mm long, smooth, globose | E. scaligera |
| → 53 |
53. Primary panicle branches not rebranched; proximal spikelets on each branch sessile or subsessile, the pedicels shorter than 0.4 mm | E. sessilispica |
53. Primary panicle branches usually with secondary branches; proximal spikelets on each branch usually pedicellate, the pedicels longer than 0.4 mm. | → 54 |
54. Spikelets 1.3-2 mm long, with 1-3 florets; lemmas 0.8-1.2 mm long | E. airoides |
54. Spikelets 2-19 mm long, with 2-22 florets; lemmas 1.2-2.4 mm long. | → 55 |
55. Spikelets 2-4.5(5) mm long. | → 56 |
56. Blades 25-60 cm long, 3-11 mm wide; lemmas 1.6-2.4 mm long; spikelets 1-1.7 mm wide; sheaths densely hirsute, with papillose-based hairs on the base, back, and collar | E. hirsuta |
56. Blades 4-22 cm long, 1-3.5 mm wide; lemmas 1.2-1.8 mm long; spikelets 0.5-1.3 mm wide; sheaths sometimes hirsute, at least partially, but the hairs never papillose-based | E. lugens |
55. Spikelets 4-19 mm long. | → 57 |
57. Spikelets with 10-22 florets; caryopses terete to laterally compressed, opaque, uniformly reddish brown | E. atrovirens |
57. Spikelets with 3-12(14) florets; caryopses dorsally compressed, translucent, greenish over the embryo. | → 58 |
58. Lemmas 1.8-3 mm long; panicles 16-35(40) cm long, (4)8-24 cm wide; blades 12-50(65) cm long; caryopses 1-1.7 mm long; ligules 0.6-1.3 mm long | E. curvula |
58. Lemmas 1.4-1.7 long; panicles 6-18 cm long, 2-8 cm wide; blades 2-12 cm long; caryopses 0.4-0.8 mm long; ligules 0.3-0.5 mm long. | → 59 |
59. Plants without woolly hairs on the base; glumes unequal; lateral lemma veins not green, inconspicuous throughout; spikelets 0.8-1.2 mm; naturalized in the southwestern United States | E. lehmanniana |
59. Plants with conspicuous, woolly hairs on the base; glumes subequal; lateral lemma veins green, conspicuous basally, obscure near the lemma apices; spikelets 1.3-2 mm wide; in the Flora region, known only from waste areas near a woolen mill in South Carolina | E. setifolia |