Epilobium saximontanum |
Epilobium torreyi |
|
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glandular willowherb, Rocky Mountain willowherb, Rocky Mountain willowweed, épilobe des rocheuses |
brook spike-primrose, brook willowherb, narrow boisduvalia, stiff spikeprimrose, Torrey's epilobium, Torrey's willowherb |
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Habit | Herbs usually with sessile, fleshy, underground turions, or sometimes thick, elongated shoots with dark, decussate scales. | Herbs with taproot. |
Stems | erect, strict, terete, 4–55 cm, simple or well branched in age, subglabrous proximally to mixed strigillose and glandular puberulent distally, with raised strigillose lines decurrent from margins of petioles. |
erect, terete, (1–)4–65 cm, usually with several erect or ascending virgate branches proximally or simple, densely villous, sometimes mixed strigillose, or sometimes subglabrous. |
Leaves | opposite proximal to inflorescence, alternate and reduced distally, often ± appressed, usually subsessile, rarely petiole 1–3 mm, often clasping; blade obovate proximally to ovate, lanceolate, or narrowly elliptic distally, 1–5.5(–6.5) × 0.4–2(–2.4) cm, base rounded or obtuse, margins low denticulate, 9–30 teeth per side, veins ± conspicuous, 3–6 per side, apex subacute, surfaces subglabrous with strigillose margins; bracts much reduced. |
opposite only in proximal pairs, distally alternate, subsessile, blade linear-lanceolate to very narrowly elliptic, 0.5–4.5 × 0.2–0.3(–0.6) cm, usually longer than internodes, base cuneate, margins subentire to sparsely serrulate, 2–5 low teeth per side, lateral veins obscure, 2–5 per side, apex acute, surfaces subglabrous proximally to densely villous and/or strigillose distally; bracts slightly reduced. |
Inflorescences | erect, sometimes nodding in bud, racemes, sometimes sparsely branched. |
erect spikes, simple or sparsely branched, ± densely villous and/or strigillose. |
Flowers | erect; buds 2–3.5 × 1.8–2.5 mm; pedicel 0–1 mm; floral tube 0.8–1.4 × 0.8–1.9 mm, ring of sparse spreading hairs at mouth inside; sepals sometimes flushed red, 1.2–3.5 × 0.6–1.4 mm, abaxial surface strigillose and sometimes mixed glandular puberulent; petals usually white, infrequently pink, 2.2–5(–7) × 1.7–3.2 mm, apical notch 0.4–1.5 mm; filaments usually cream, rarely light pink, those of longer stamens 2–3.5 mm, those of shorter ones 1–2 mm; anthers cream to light yellow, 0.3–0.8 × 0.3–0.5 mm; ovary 9–30 mm, densely strigillose and glandular puberulent; style cream or yellow, 1.6–2.8 mm, stigma usually narrowly to broadly clavate, rarely subcapitate, 1–3 × 0.8–2 mm, surrounded by at least longer anthers. |
erect, often initiating in most proximal nodes, usually cleistogamous; buds 0.8–1.5 × 0.5–1 mm; floral tube 0.4–1 × 0.6–1.2 mm, ring of lax hairs near base inside; sepals 0.7–2 × 0.4–0.8 mm; petals pink with darker veins or white, 1.2–3.2 × 0.9–1.8 mm, apical notch 0.4–1 mm; filaments pale pink, those of longer stamens 0.4–1.8 mm, those of shorter ones 0.3–1 mm; anthers 0.3–0.6 × 0.3–0.4 mm, apiculate; ovary 3–6 mm, densely pubescent; style pale pink, 1.2–2.8 mm, stigma clavate to subcapitate or irregularly 4-lobed, 0.5–1.2 ×0.3–0.6 mm, surrounded by longer anthers. |
Capsules | 30–55(–70) mm, surfaces mixed strigillose and glandular puberulent; usually subsessile, rarely pedicel 1–5 mm, often appressed to stem. |
cylindrical to subfusiform, terete to slightly 4-angled, (6–)8–14 × 1.1–2 mm, beak 2–3 mm, central column disintegrating, villous; sessile. |
Seeds | very narrowly obovoid, 1–1.6(–1.8) × 0.4–0.6 mm, chalazal collar 0.1–0.2 mm, light brown or gray, surface rugose to papillose; coma usually readily detached, white, 3–9 mm. |
3–6 per locule, irregularly angular-oblong or fusiform, 0.9–1.6 × 0.4–0.9 mm, chalazal collar absent, brown, surface irregularly reticulate. |
2n | = 36. |
= 18. |
Epilobium saximontanum |
Epilobium torreyi |
|
Phenology | Flowering Jul–Sep. | Flowering May–Aug. |
Habitat | Montane semi-shaded stream banks, damp meadows, mossy seeps, wet slatey cliffs, disturbed or seasonally damp areas. | Moist places along stream banks, seasonal streambeds, seeps, and roadside ditches, often in gravelly red or granite soil. |
Elevation | 0–3700 m. (0–12100 ft.) | 0–2600 m. (0–8500 ft.) |
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WY; AB; BC; MB; NF; ON; QC
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CA; ID; NV; OR; WA; BC
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Discussion | Epilobium saximontanum is morphologically similar to E. ciliatum (especially subsp. glandulosum) with which it also shares the AA chromosome arrangement. However, in addition to its fleshy compact turions, it very characteristically has notably appressed capsules, unlike most other species in the genus, and a notably strict habit. The distribution of Epilobium saximontanum is unusual; it includes the Rocky Mountain region, only barely reaching the high southern Sierra Nevada, disjunct to the Black Hills of South Dakota, and more widely in eastern Canada, from the shores of Hudson Bay to Newfoundland. Specimens are fairly uniform across this wide and rather discontinuous range, although locally they show some variability, possibly due to hybridization with any of several species that may be sympatric with it. H. Lewis and D. M. Moore (1962) reported hybrids between E. saximontanum (cited as E. brevistylum) and E. ciliatum subsp. ciliatum (cited as E. adenocaulon) from Colorado, and herbarium specimens with E. saximontanum and apparent hybrids are not uncommon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Epilobium torreyi is an annual species that has a unique gametic chromosome number of n = 9, the lowest found in the genus. Its morphological similarity to Epilobium pallidum (n = 10) suggested to P. H. Raven and D. M. Moore (1965) that E. torreyi may be an aneuploid derivative of E. pallidum or a close relative. No natural hybrids among species of sect. Pachydium have been reported, but in fact such plants might be difficult to detect. S. R. Seavey (1992) was able to form hybrids, but they were highly sterile. Nevertheless, hybridization between species with n = 9 and n = 10, followed by polyploidization, may well have given rise to the South American species of this section, Epilobium subdentatum (n = 19). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobium | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Pachydium |
Sibling taxa | ||
Synonyms | E. adenocaulon subsp. rubescens, E. drummondii, E. drummondii var. latiusculum, E. latiusculum, E. ovatifolium, E. rubescens, E. scalare, E. stramineum | Oenothera torreyi, Boisduvalia parviflora, B. stricta, B. torreyi, Gayophytum strictum, Oenothera densiflora var. tenella |
Name authority | Haussknecht: Oesterr. Bot. Z. 29: 119. (1879) | (S. Watson) Hoch & P. H. Raven: Phytologia 73: 458. (1993) |
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