Epilobium saximontanum |
Epilobium canum |
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glandular willowherb, Rocky Mountain willowherb, Rocky Mountain willowweed, épilobe des rocheuses |
California fire chalice, California fuchsia, firechalice, hummingbird trumpet, zauschneria |
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Habit | Herbs usually with sessile, fleshy, underground turions, or sometimes thick, elongated shoots with dark, decussate scales. | Herbs suffruticose or not, with basal shoots from ± woody caudex, often decussate scales at base. | ||||||||
Stems | erect, strict, terete, 4–55 cm, simple or well branched in age, subglabrous proximally to mixed strigillose and glandular puberulent distally, with raised strigillose lines decurrent from margins of petioles. |
erect to ascending, often clumped but not matted, green or gray-green, terete, 10–110(–120) cm, usually well-branched throughout, sometimes simple, strigillose and/or long-villous, usually mixed glandular puberulent distally, rarely glabrate. |
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Leaves | opposite proximal to inflorescence, alternate and reduced distally, often ± appressed, usually subsessile, rarely petiole 1–3 mm, often clasping; blade obovate proximally to ovate, lanceolate, or narrowly elliptic distally, 1–5.5(–6.5) × 0.4–2(–2.4) cm, base rounded or obtuse, margins low denticulate, 9–30 teeth per side, veins ± conspicuous, 3–6 per side, apex subacute, surfaces subglabrous with strigillose margins; bracts much reduced. |
± densely spaced, alternate and often fasciculate distally, subsessile, blade grayish green or green to silvery-canescent, usually narrowly linear to lanceolate or elliptic to ovate, rarely orbiculate, 0.6–5(–6) × 0.1–2.5 cm, base cuneate to attenuate, margins subentire to sharply toothed, 3–15 teeth per side, veins inconspicuous or prominent, 3–7 per side, apex acute, sometimes with caducous dark mucro, surfaces usually ± densely strigillose, sometimes mixed villous and/or glandular puberulent, rarely glabrate; bracts much smaller and narrower. |
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Inflorescences | erect, sometimes nodding in bud, racemes, sometimes sparsely branched. |
erect spikes or racemes, loose to congested, often branched, glandular puberulent and sometimes mixed strigillose or villous. |
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Flowers | erect; buds 2–3.5 × 1.8–2.5 mm; pedicel 0–1 mm; floral tube 0.8–1.4 × 0.8–1.9 mm, ring of sparse spreading hairs at mouth inside; sepals sometimes flushed red, 1.2–3.5 × 0.6–1.4 mm, abaxial surface strigillose and sometimes mixed glandular puberulent; petals usually white, infrequently pink, 2.2–5(–7) × 1.7–3.2 mm, apical notch 0.4–1.5 mm; filaments usually cream, rarely light pink, those of longer stamens 2–3.5 mm, those of shorter ones 1–2 mm; anthers cream to light yellow, 0.3–0.8 × 0.3–0.5 mm; ovary 9–30 mm, densely strigillose and glandular puberulent; style cream or yellow, 1.6–2.8 mm, stigma usually narrowly to broadly clavate, rarely subcapitate, 1–3 × 0.8–2 mm, surrounded by at least longer anthers. |
buds 11–18 × 4–6 mm, subsessile or pedicels 1–2 mm; floral tube same color as petals, 16–32 × 5–8 mm, base slightly bulbous, ring of 8 irregular scales at base of filaments 4–6.5 mm from base inside; sepals same color as petals, 7–15 × 3.5–5 mm, abaxial surface densely pubescent; petals usually orange-red, very rarely white, obcordate, 8–17 × 5–9.5 mm, apical notch 2–3 mm; filaments light orange-red to white, those of longer stamens 12.5–32 mm, those of shorter ones 10–25 mm; anthers 2.7–4x 0.8–1.2 mm, apiculate; ovary 8–15 mm, glandular puberulent, often mixed villous; style light orange-red, 42–65 mm, glabrous, stigma 4-lobed, 1–1.4 × 2.4–3 mm, exserted 8–15 mm beyond anthers. |
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Capsules | 30–55(–70) mm, surfaces mixed strigillose and glandular puberulent; usually subsessile, rarely pedicel 1–5 mm, often appressed to stem. |
straight or ± curved-ascending, 15–35 mm, sometimes beaked, surfaces glandular puberulent and strigillose; subsessile or pedicel 0–3 mm. |
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Seeds | very narrowly obovoid, 1–1.6(–1.8) × 0.4–0.6 mm, chalazal collar 0.1–0.2 mm, light brown or gray, surface rugose to papillose; coma usually readily detached, white, 3–9 mm. |
broadly to narrowly obovoid, with constriction 0.6–0.8 mm from micropylar end, 1.5–2.6 × 0.9–1.3 mm, chalazal collar inconspicuous, light brown, surface low papillose; coma easily detached, dingy white, 5.5–7 mm. |
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2n | = 36. |
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Epilobium saximontanum |
Epilobium canum |
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Phenology | Flowering Jul–Sep. | |||||||||
Habitat | Montane semi-shaded stream banks, damp meadows, mossy seeps, wet slatey cliffs, disturbed or seasonally damp areas. | |||||||||
Elevation | 0–3700 m. (0–12100 ft.) | |||||||||
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WY; AB; BC; MB; NF; ON; QC
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sw United States; nw Mexico
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Discussion | Epilobium saximontanum is morphologically similar to E. ciliatum (especially subsp. glandulosum) with which it also shares the AA chromosome arrangement. However, in addition to its fleshy compact turions, it very characteristically has notably appressed capsules, unlike most other species in the genus, and a notably strict habit. The distribution of Epilobium saximontanum is unusual; it includes the Rocky Mountain region, only barely reaching the high southern Sierra Nevada, disjunct to the Black Hills of South Dakota, and more widely in eastern Canada, from the shores of Hudson Bay to Newfoundland. Specimens are fairly uniform across this wide and rather discontinuous range, although locally they show some variability, possibly due to hybridization with any of several species that may be sympatric with it. H. Lewis and D. M. Moore (1962) reported hybrids between E. saximontanum (cited as E. brevistylum) and E. ciliatum subsp. ciliatum (cited as E. adenocaulon) from Colorado, and herbarium specimens with E. saximontanum and apparent hybrids are not uncommon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). This treatment recognizes three self-compatible but highly outcrossing subspecies marked by distinct but sometimes intergrading morphology and overlapping geographical ranges. R. N. Bowman and P. C. Hoch (1979) agreed with the treatment of Epilobium canum subsp. garrettii (n = 15) and subsp. latifolium (n = 30) by P. H. Raven (1976), but considering the complex intergrading patterns of variation involving the rest of this species, they combined the two remaining tetraploid subspecies recognized by Raven (subspp. angustifolium and mexicanum) with the remaining diploid subspecies into a single polyploid subsp. canum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobium | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Zauschneria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | E. adenocaulon subsp. rubescens, E. drummondii, E. drummondii var. latiusculum, E. latiusculum, E. ovatifolium, E. rubescens, E. scalare, E. stramineum | Zauschneria cana | ||||||||
Name authority | Haussknecht: Oesterr. Bot. Z. 29: 119. (1879) | (Greene) P. H. Raven: Ann. Missouri Bot. Gard. 63: 335. (1977) | ||||||||
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