Epilobium leptophyllum |
Epilobium pallidum |
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bog willow-herb, linearleaf willowweed, narrow-leaf willowherb, slender leaf willowherb, épilobe leptophylle |
largeflower spike-primrose, pale boisduvalia |
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Habit | Herbs with threadlike, nearly leafless epigeous stolons terminating in compact, fleshy turions 3–8 × 2–4 mm. | Herbs with taproot. |
Stems | erect, simple to loosely clustered, terete, 15–95 cm, simple to well branched, densely strigillose, often mixed glandular puberulent on inflorescence, rarely with faint strigillose lines decurrent from margins of petioles. |
terete, 4.5–55 cm, simple or with proximal ascending branches and/or distal branches, ± densely villous and/or strigillose, mixed glandular puberulent distally or glabrescent. |
Leaves | opposite proximally, usually alternate, rarely fasciculate distally, subsessile; blade linear to very narrowly elliptic or sublanceolate, 2–7.5 × 0.1–0.7 cm, usually longer than internodes, base rounded to subcuneate, margins subentire, 4–7 inconspicuous teeth per side, sometimes revolute, lateral veins inconspicuous, apex obtuse proximally to acute distally, both surfaces densely strigillose, increasing distally; bracts not much reduced. |
opposite only in proximal pairs, alternate and crowded distally, subsessile, blade narrowly elliptic or lanceolate, 1.2–5 × 0.2–1.2 cm, usually longer than internodes, base cuneate to attenuate, margins subentire or sparsely serrulate, 3–5 teeth per side, veins inconspicuous, 2–7 per side, apex acute, surfaces villous; bracts same size and shape as leaves. |
Inflorescences | erect racemes, densely strigillose, often mixed sparsely glandular puberulent. |
erect spikes, often congested proximally, sometimes nodding in bud, simple, or sometimes sparsely branched, ± densely mixed villous, strigillose, and glandular puberulent. |
Flowers | erect; buds 3–5 × 1.5–2.5 mm; pedicel 5–12 mm; floral tube 0.8–1.5 × 1.2–1.8 mm, ring of spreading hairs at mouth inside; sepals 2.5–4.5 × 0.9–1.3 mm, abaxial surface strigillose; petals obcordate, white to light pink, 3.5–7 × 1.6–4 mm, apical notch 1–1.8 mm; filaments white or cream, those of longer stamens 0.8–3.5 mm, those of shorter ones 0.6–2.5 mm; anthers cream, 0.5–0.9 × 0.4–0.6 mm; ovary 12–18 mm, densely strigillose, sometimes mixed glandular puberulent; style cream, 2–3.8 mm, stigma narrowly clavate, entire, 1–1.8 × 0.5–1.2 mm, usually surrounded by, rarely exserted beyond, anthers. |
erect, mostly chasmogamous, sometimes initiating at second or third proximal node; buds 3–5 × 2.5–3.5 mm, apex blunt; floral tube 1.5–3 × 1.1–2.8 mm, ring of lax hairs near base inside; sepals erect, (1.5–)2–6.2 × (0.5–)1–2.2 mm, densely villous and glandular pubescent abaxially; petals rose-purple, (2.8–)4.2–9.8 × (2.2–)3.5–5.3 mm, apical notch 1.2–3.5 mm; filaments rose-purple, those of longer stamens 2.4–6.5 mm, those of shorter ones 0.9–4.3 mm; anthers 0.6–1.6 × 0.2–0.8 mm; ovary 3.5–9 mm, densely pubescent; style pink, 3.3–9.2 mm, stigma 4-lobed, ± subentire in smaller flowers, 0.7–1.4 × 0.5–1.3 mm, usually surrounded by, rarely exserted beyond, longer anthers. |
Capsules | straight, narrowly cylindrical, 35–80 mm, surfaces densely strigillose; pedicel 10–35 mm. |
narrowly fusiform, (10–)14–21 × 1–2.1 mm, beak 2–5 mm, ribs usually broad and prominent, septa and central column disintegrating with age, surfaces ± densely villous; subsessile. |
Seeds | narrowly fusiform to narrowly obovoid, 1.5–2.2 ×0.5–0.7 mm, chalazal collar 0.1–0.2 mm, ± pronounced, surface papillose; coma persistent, dingy white, 6–8 mm. |
8–12 per capsule, initially in 4 locules, at maturity pushed into 2 rows proximally, 1 overlapping row distally, irregularly angular-fusiform, 1.8–2.3x 0.7–0.9 mm, chalazal collar absent, surface irregularly reticulate. |
2n | = 36. |
= 20. |
Epilobium leptophyllum |
Epilobium pallidum |
|
Phenology | Flowering Jun–Sep. | Flowering May–Aug(–Sep). |
Habitat | Marshy ground, bogs, fens, low thickets, seepage areas, damp pastures. | Stream banks and washes in vernally moist areas in shrubland or lower forested regions. |
Elevation | 0–1000(–2900) m. (0–3300(–9500) ft.) | 60–2100 m. (200–6900 ft.) |
Distribution |
AK; CA; CO; CT; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; SPM
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CA; ID; OR
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Discussion | The range of Epilobium leptophyllum overlaps with that of the related E. palustre, but the former is less common to the north and more common south into the midwestern United States, and absent only from most of the southern tier of states. It is also relatively uncommon in the western United States and Canada. Judging by the number of herbarium sheets that also include E. palustre, E. densum, and even E. coloratum, it sometimes occurs in sympatry with those species and may rarely hybridize with them, based on plants with intermediate morphology and/or sterile fruits. Fernald described Epilobium nesophilum from the Magdalen Islands (Quebec), and especially Newfoundland, first as a variety of E. densum (1918), then as a separate species (1925). Epilobium rosmarinifolium Pursh 1813, an illegitimate name (not Haenke 1788), pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
P. H. Raven and D. M. Moore (1965) observed that smaller-flowered individuals of this species were sometimes difficult to separate from Epilobium torreyi (reported as Boisduvalia stricta), especially in the absence of mature fruits. The capsules of E. pallidum are usually longer and thinner than those of related species and the internal structure is highly distinctive. The stigmas of larger-flowered plants often are exserted beyond the anthers, increasing the chances for outcrossing. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobium | Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Pachydium |
Sibling taxa | ||
Synonyms | E. densum var. nesophilum, E. nesophilum, E. oliganthum var. gracile, E. palustre var. gracile, E. squamatum, E. tenellum | Boisduvalia pallida, B. macrantha |
Name authority | Rafinesque: Précis Découv. Somiol., 41. (1814) | (Eastwood) Hoch & P. H. Raven: Phytologia 73: 458. (1993) |
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