Epilobium hirsutum
Epilobium montanum
codlins-and-cream, fiddle grass, fiddle grass willowherb, great or great hairy willowherb, great willowherb, hairy willlowherb, hairy willow-herb, épilobe hirsute
broad-leaf willow-herb, tooth willow-herb, willowherb
erect to ascending, often clumped, terete, 25–120(–250) cm, unusually thick, 3–9 mm diam., well branched mainly in distal 1/2, densely long-villous throughout, usually mixed glandular puberulent distally, rarely sparsely villous or densely white-tomentose.
erect, terete, (5–)20–95 cm, often well branched, especially distally, densely strigillose proximal to inflorescence, without decurrent lines, mixed strigillose and glandular puberulent distally.
opposite proximal to inflorescence, alternate distally, sessile and ± clasping stem;
blade elliptic-lanceolate to narrowly obovate or elliptic, 4–12(–23) × 0.3–4(–5) cm, base cuneate to attenuate, margins serrulate, 15–50 teeth per side, veins 6–9 per side, apex acute to acuminate or obtuse proximally, surfaces ± densely villous;
bracts moderately reduced.
opposite proximal to inflorescence, alternate distally, petiole 1–6 mm;
blade dark green, narrowly ovate to ovate, 3–8 × 1.5–3.4 cm, subequal to internodes, base broadly cuneate to truncate, margins irregularly serrulate with 20–30 teeth per side, veins prominent, 3–5 per side, apex acuminate to acute, surfaces strigillose, especially along veins and margins;
bracts much reduced.
erect racemes or panicles, usually densely villous and glandular puberulent, rarely tomentose.
nodding to erect, racemes or loose panicles, strigillose and glandular puberulent.
erect;
buds 5–9 × 1.8–4.5 mm, sometimes beaked;
pedicel 3–11 mm;
floral tube 1.3–2.9 × 2.2–4 mm, conspicuous ring of spreading hairs near mouth inside;
sepals oblong-linear, often keeled, 6–12 × 1–3 mm, abaxial surface densely pubescent;
petals bright pink to rose-purple, rarely white, broadly obcordate, 9–20 × 7–15 mm, apical notch 1–3 mm;
filaments white or pink, those of longer stamens 5–10 mm, those of shorter ones 2.5–6 mm;
anthers cream, 1.5–3 × 0.6–1.2 mm;
ovary 15–34 mm, densely villous and glandular puberulent;
style white or pink, 5–12 mm, usually glabrous, stigma deeply 4-lobed, 1.8–2.2 × 3–5.5 mm, lobes recurved or spreading, exserted beyond anthers.
erect;
buds 4–5x 1.5–3 mm;
pedicel 2–18 mm;
floral tube 1.2–2 × 1.5–2.5 mm, conspicuous ring of spreading hairs at mouth inside;
sepals often flushed red, keeled, 5–6.5 × 1.2–1.7 mm, abaxial surface strigillose and glandular puberulent;
petals rose-purple, obcordate, 7.5–10 × 4–5.5 mm, apical notch 2.7–4.2 mm;
filaments light pink, those of longer stamens 3.5–5.5 mm, those of shorter ones 1.5–2.4 mm;
anthers yellow, 1–1.2 × 0.5–0.6 mm;
ovary 20–30 mm, strigillose and glandular puberulent;
style rose-purple to white, 3.5–7.5 mm, glabrous or with scattered long hairs near base, stigma deeply 4-lobed, 1.5–2 × 2.5–4 mm, lobes 1.2–1.8 mm, exserted beyond or, sometimes, surrounded by anthers.
often flushed purple, 25–90 mm, surfaces usually densely villous and glandular puberulent, rarely glabrescent;
pedicel 5–20 mm.
40–80 mm, surfaces strigillose and glandular puberulent;
pedicel 8–20 mm.
narrowly obovoid, 0.8–1.2 × 0.3–0.6 mm, chalazal collar inconspicuous, dark brown, surface coarsely papillose;
coma easily detached, tawny or dull white, 7–10 mm.
obovoid, 1–1.2 × 0.4–0.5 mm, chalazal collar inconspicuous, 0.6–0.8 mm wide, brown, surface coarsely papillose;
coma readily detached, dull white, 5–6 mm.
= 36.
= 36.
Epilobium hirsutum
Epilobium montanum
Epilobium hirsutum is very widespread in cool temperate Eurasia and montane regions, occurring throughout Europe except in the far north (P. H. Raven 1968), through the Caucasus and central Asia (E. I. Steinberg 1949) to Nepal (Raven 1962), China (Chen C. J. et al. 1992), and Japan (A. W. Lievens and P. C. Hoch 1999). It occurs as well along the Mediterranean coast of Africa, through East Africa to southern Africa, and in the Canary and Cape Verde Islands (Raven 1967).
Epilobium hirsutum exceeds almost all other species of the genus in stature, so its size, very large flowers, and densely villous aspect make it easy to identify. R. L. Stuckey (1970) provided a detailed account of the introduction and spread of E. hirsutum in North America, noting the earliest known collection (July 1829) was from Newport, Rhode Island. Most early collections appeared in waste areas, particularly near harbor ballast piles, although some may have been grown in gardens. By the 1890s this species was well established along the Atlantic coastal region from New Jersey and Philadelphia through New England, and around Niagara Falls in the Great Lakes region. During the twentieth century, E. hirsutum spread extensively in southern Ontario and Quebec, south along the Atlantic coast to Maryland, and to all of the states along the southern shores of the Great Lakes, most recently including Wisconsin (1970), and Indiana (1972). It occurs in much the same habitat as that of another, more widely publicized invader, Lythrum salicaria, and sometimes is recorded as a companion species. The earliest known collection in western North America was made in 1933 in Bingen (Klickitat County), Washington. Whether from that introduction or others, E. hirsutum is now naturalized and widespread in the Pacific Northwest. It also was reported recently from the Denver region in Colorado and near Midway in Utah.
Epilobium grandiflorum F. H. Wiggers and E. grandiflorum Allioni are illegitimate names that pertain here.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Epilobium montanum is native and widely distributed throughout Europe and in Asia at least as far east as the Ural Mountains and the Caucasus, and in Japan (A. W. Lievens and P. C. Hoch 1999). It also is questionably naturalized on Campbell Island, New Zealand (P. H. Raven and T. E. Raven 1976). Although E. montanum was first collected in 1894 in the vicinity of St. John’s, Newfoundland, and clearly naturalized in that area, it does not appear to have spread appreciably in eastern North America, unlike European adventives such as E. hirsutum or E. parviflorum. The collection by F. C. Seymour in 1971 in Maine may be an independent introduction. Collections of E. montanum in the University of British Columbia Botanical Garden in Vancouver appeared to have been ephemeral, but more recent collections in ruderal areas in that region suggest that it has naturalized there. Like other European introductions, E. montanum has the BB chromosome arrangement (S. R. Seavey and P. H. Raven 1977).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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