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codlins-and-cream, fiddle grass, fiddle grass willowherb, great or great hairy willowherb, great willowherb, hairy willlowherb, hairy willow-herb, épilobe hirsute

glandular willowherb, Hall's willowherb, Sierra willowherb

Habit Herbs usually robust and rank, sometimes woody near base, with thick, ropelike stolons to 1 m with scattered cataphylls and, often, terminal leafy rosette. Herbs with small, 3–6 mm, round or oblong, compact turions 1–5 cm below ground.
Stems

erect to ascending, often clumped, terete, 25–120(–250) cm, unusually thick, 3–9 mm diam., well branched mainly in distal 1/2, densely long-villous throughout, usually mixed glandular puberulent distally, rarely sparsely villous or densely white-tomentose.

strict, erect, terete, 2–50(–60) cm, rarely branched only in larger plants, subglabrous proximal to inflorescence except for raised strigillose lines decurrent from margins of petioles, or sometimes ± densely long-villous throughout with inconspicuous decurrent lines.

Leaves

opposite proximal to inflorescence, alternate distally, sessile and ± clasping stem;

blade elliptic-lanceolate to narrowly obovate or elliptic, 4–12(–23) × 0.3–4(–5) cm, base cuneate to attenuate, margins serrulate, 15–50 teeth per side, veins 6–9 per side, apex acute to acuminate or obtuse proximally, surfaces ± densely villous;

bracts moderately reduced.

opposite proximally, alternate on inflorescence, subsessile or proximally with petioles 1–1.5 mm;

blade ovate proximally to lanceolate or narrowly elliptic distally, 0.5–4.7 × 0.2–1.4 cm, base rounded to cuneate, margins subentire proximally to denticulate distally, 8–20 teeth per side, veins inconspicuous, 3–6 per side, apex obtuse to subacute, surfaces mostly glabrous with strigillose margins;

bracts much reduced.

Inflorescences

erect racemes or panicles, usually densely villous and glandular puberulent, rarely tomentose.

usually nodding in bud, erect later, open racemes, sometimes congested, usually mixed strigillose and glandular puberulent, rarely also mixed villous, or rarely subglabrous.

Flowers

erect;

buds 5–9 × 1.8–4.5 mm, sometimes beaked;

pedicel 3–11 mm;

floral tube 1.3–2.9 × 2.2–4 mm, conspicuous ring of spreading hairs near mouth inside;

sepals oblong-linear, often keeled, 6–12 × 1–3 mm, abaxial surface densely pubescent;

petals bright pink to rose-purple, rarely white, broadly obcordate, 9–20 × 7–15 mm, apical notch 1–3 mm;

filaments white or pink, those of longer stamens 5–10 mm, those of shorter ones 2.5–6 mm;

anthers cream, 1.5–3 × 0.6–1.2 mm;

ovary 15–34 mm, densely villous and glandular puberulent;

style white or pink, 5–12 mm, usually glabrous, stigma deeply 4-lobed, 1.8–2.2 × 3–5.5 mm, lobes recurved or spreading, exserted beyond anthers.

erect;

buds 2–5 × 1–2 mm;

pedicel 3–8 mm;

floral tube 0.5–1.7 × 0.8–1.6 mm, with slightly raised ring of spreading hairs at mouth inside;

sepals green, 1.2–2.8 × 0.5–1 mm, abaxial surface subglabrous or sparsely glandular puberulent;

petals white, often fading pink, 1.6–5.5 × 1.2–3 mm, apical notch 0.3–1.2 mm;

filaments white or cream, those of longer stamens 0.6–2.5 mm, those of shorter ones 0.4–1.5 mm;

anthers cream, 0.2–0.9 × 0.2–0.5 mm;

ovary 10–14 mm, strigillose and glandular puberulent or subglabrous;

style cream, 0.8–5 mm, stigma clavate, entire, 0.4–1.2 × 0.3–0.7 mm, usually surrounded by, rarely exserted beyond, anthers.

Capsules

often flushed purple, 25–90 mm, surfaces usually densely villous and glandular puberulent, rarely glabrescent;

pedicel 5–20 mm.

very narrowly cylindrical, (15–)24–60 mm, surfaces usually subglabrous to mixed strigillose and glandular puberulent, rarely sparsely villous;

pedicel 8–40 mm.

Seeds

narrowly obovoid, 0.8–1.2 × 0.3–0.6 mm, chalazal collar inconspicuous, dark brown, surface coarsely papillose;

coma easily detached, tawny or dull white, 7–10 mm.

narrowly fusiform to narrowly obovoid, 1.1–1.6 × 0.4–0.6 mm, chalazal collar ± conspicuous, 0.05–0.2 mm, light brown, surface papillose;

coma easily detached, white, 3–6 mm.

2n

= 36.

= 36.

Epilobium hirsutum

Epilobium hallianum

Phenology Flowering Jun–Sep. Flowering Jun–Sep.
Habitat Low wet areas along streams, rivers, ponds, and lakes, roadside ditches, along railroad tracks, marshes and swampy areas. Semi-shaded stream banks, wet grassy slopes or meadows, bogs, seasonally wet sites, vernal pools.
Elevation 0–150[–3000] m. (0–500[–9800] ft.) 100–3700 m. (300–12100 ft.)
Distribution
from FNA
CO; CT; IL; IN; KY; MA; MD; ME; MI; NH; NJ; NY; OH; OR; PA; RI; UT; VT; WA; WI; WV; BC; NB; NS; ON; PE; QC; Eurasia; Africa [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Epilobium hirsutum is very widespread in cool temperate Eurasia and montane regions, occurring throughout Europe except in the far north (P. H. Raven 1968), through the Caucasus and central Asia (E. I. Steinberg 1949) to Nepal (Raven 1962), China (Chen C. J. et al. 1992), and Japan (A. W. Lievens and P. C. Hoch 1999). It occurs as well along the Mediterranean coast of Africa, through East Africa to southern Africa, and in the Canary and Cape Verde Islands (Raven 1967).

Epilobium hirsutum exceeds almost all other species of the genus in stature, so its size, very large flowers, and densely villous aspect make it easy to identify. R. L. Stuckey (1970) provided a detailed account of the introduction and spread of E. hirsutum in North America, noting the earliest known collection (July 1829) was from Newport, Rhode Island. Most early collections appeared in waste areas, particularly near harbor ballast piles, although some may have been grown in gardens. By the 1890s this species was well established along the Atlantic coastal region from New Jersey and Philadelphia through New England, and around Niagara Falls in the Great Lakes region. During the twentieth century, E. hirsutum spread extensively in southern Ontario and Quebec, south along the Atlantic coast to Maryland, and to all of the states along the southern shores of the Great Lakes, most recently including Wisconsin (1970), and Indiana (1972). It occurs in much the same habitat as that of another, more widely publicized invader, Lythrum salicaria, and sometimes is recorded as a companion species. The earliest known collection in western North America was made in 1933 in Bingen (Klickitat County), Washington. Whether from that introduction or others, E. hirsutum is now naturalized and widespread in the Pacific Northwest. It also was reported recently from the Denver region in Colorado and near Midway in Utah.

Epilobium grandiflorum F. H. Wiggers and E. grandiflorum Allioni are illegitimate names that pertain here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Epilobium hallianum has condensed fleshy turions and generally strict habit, suggesting an affinity with E. ciliatum or E. saximontanum, both with the AA arrangement. However, E. hallianum has the BB arrangement (S. R. Seavey and P. H. Raven 1978) and apparently more distant relationship to those other species.

In different parts of its geographical range Epilobium hallianum shows considerable morphological variability, especially in leaf shape and margins, and in type and pattern of stem vestiture, including plants in the southern part of its range with densely villous stems that have been treated as E. ursinum. In part because of this variability and in part because the very characteristic condensed turions are easily lost during collection and/or often overlooked, E. hallianum is frequently misidentified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobium Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobium
Sibling taxa
E. anagallidifolium, E. arcticum, E. brachycarpum, E. campestre, E. canum, E. ciliatum, E. clavatum, E. cleistogamum, E. coloratum, E. davuricum, E. densiflorum, E. densum, E. foliosum, E. glaberrimum, E. hallianum, E. hornemannii, E. howellii, E. lactiflorum, E. leptocarpum, E. leptophyllum, E. luteum, E. minutum, E. mirabile, E. montanum, E. nevadense, E. nivium, E. obcordatum, E. obscurum, E. oreganum, E. oregonense, E. pallidum, E. palustre, E. parviflorum, E. rigidum, E. saximontanum, E. septentrionale, E. siskiyouense, E. smithii, E. suffruticosum, E. torreyi
E. anagallidifolium, E. arcticum, E. brachycarpum, E. campestre, E. canum, E. ciliatum, E. clavatum, E. cleistogamum, E. coloratum, E. davuricum, E. densiflorum, E. densum, E. foliosum, E. glaberrimum, E. hirsutum, E. hornemannii, E. howellii, E. lactiflorum, E. leptocarpum, E. leptophyllum, E. luteum, E. minutum, E. mirabile, E. montanum, E. nevadense, E. nivium, E. obcordatum, E. obscurum, E. oreganum, E. oregonense, E. pallidum, E. palustre, E. parviflorum, E. rigidum, E. saximontanum, E. septentrionale, E. siskiyouense, E. smithii, E. suffruticosum, E. torreyi
Synonyms Chamaenerion hirsutum, E. amplexicaule, E. aquaticum, E. hirsutum var. villosum, E. villosum E. brevistylum var. pringleanum, E. brevistylum var. subfalcatum, E. brevistylum var. tenue, E. brevistylum var. ursinum, E. delicatum var. tenue, E. glandulosum var. tenue, E. pringleanum, E. pringleanum var. tenue, E. ursinum, E. ursinum var. subfalcatum
Name authority Linnaeus: Sp. Pl. 1: 347. (1753) Haussknecht: Monogr. Epilobium, 261. (1884) — (as halleanum)
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