Epilobium hirsutum
Epilobium coloratum
codlins-and-cream, fiddle grass, fiddle grass willowherb, great or great hairy willowherb, great willowherb, hairy willlowherb, hairy willow-herb, épilobe hirsute
eastern willow-herb, purple-leaf willowherb
erect to ascending, often clumped, terete, 25–120(–250) cm, unusually thick, 3–9 mm diam., well branched mainly in distal 1/2, densely long-villous throughout, usually mixed glandular puberulent distally, rarely sparsely villous or densely white-tomentose.
erect, subterete, (20–)40–85(–120) cm, freely branched distally, subglabrous proximal to inflorescence, often with raised strigillose lines decurrent from petioles, densely strigillose distally.
opposite proximal to inflorescence, alternate distally, sessile and ± clasping stem;
blade elliptic-lanceolate to narrowly obovate or elliptic, 4–12(–23) × 0.3–4(–5) cm, base cuneate to attenuate, margins serrulate, 15–50 teeth per side, veins 6–9 per side, apex acute to acuminate or obtuse proximally, surfaces ± densely villous;
bracts moderately reduced.
opposite proximal to inflorescence, alternate distally, petiole 4–10 mm;
blade narrowly lanceolate to lanceolate or narrowly elliptic, 3–10(–15) × 0.5–3 cm, subequal to internodes, base rounded to cuneate, margins sharply and irregularly serrulate, 30–75 teeth per side, veins prominent, often raised abaxially, 10–25 per side, apex acute or acuminate, surfaces subglabrous with strigillose margins and abaxial veins;
bracts abruptly reduced.
erect racemes or panicles, usually densely villous and glandular puberulent, rarely tomentose.
usually upright panicles, sometimes corymbiform, rarely racemes, densely strigillose.
erect;
buds 5–9 × 1.8–4.5 mm, sometimes beaked;
pedicel 3–11 mm;
floral tube 1.3–2.9 × 2.2–4 mm, conspicuous ring of spreading hairs near mouth inside;
sepals oblong-linear, often keeled, 6–12 × 1–3 mm, abaxial surface densely pubescent;
petals bright pink to rose-purple, rarely white, broadly obcordate, 9–20 × 7–15 mm, apical notch 1–3 mm;
filaments white or pink, those of longer stamens 5–10 mm, those of shorter ones 2.5–6 mm;
anthers cream, 1.5–3 × 0.6–1.2 mm;
ovary 15–34 mm, densely villous and glandular puberulent;
style white or pink, 5–12 mm, usually glabrous, stigma deeply 4-lobed, 1.8–2.2 × 3–5.5 mm, lobes recurved or spreading, exserted beyond anthers.
erect;
buds 2–3 × 1.5–2.5 mm;
pedicel 5–10 mm;
floral tube 0.3–0.6 × 0.3–0.5 mm, raised ring of sparse spreading hairs at mouth inside;
sepals 1.3–3.2 × 0.5–1.5 mm, abaxial surface strigillose;
petals white, 2.5–5.5 × 2–3.8 mm, apical notch 0.5–1 mm;
filaments white, those of longer stamens 1.8–2.5 mm, those of shorter ones 1.5–2 mm;
anthers pale yellow, 0.3–0.4 × 0.2–0.4 mm;
ovary 15–30 mm, ± densely strigillose;
style erect, white, 1.5–2.8 mm, glabrous, stigma cylindrical to subcapitate, entire, 1–1.5 × 0.8–1.2 mm, surrounded by longer anthers.
often flushed purple, 25–90 mm, surfaces usually densely villous and glandular puberulent, rarely glabrescent;
pedicel 5–20 mm.
40–65 mm, surfaces strigillose;
pedicel 8–12 mm.
narrowly obovoid, 0.8–1.2 × 0.3–0.6 mm, chalazal collar inconspicuous, dark brown, surface coarsely papillose;
coma easily detached, tawny or dull white, 7–10 mm.
narrowly oblanceoloid, 1.2–1.7 × 0.3–0.5 mm, abruptly rounded with very short chalazal neck, brownish gray, surface evenly papillose;
coma not easily detached, cinnamon red, 8–12 mm.
= 36.
= 36.
Epilobium hirsutum
Epilobium coloratum
Epilobium hirsutum is very widespread in cool temperate Eurasia and montane regions, occurring throughout Europe except in the far north (P. H. Raven 1968), through the Caucasus and central Asia (E. I. Steinberg 1949) to Nepal (Raven 1962), China (Chen C. J. et al. 1992), and Japan (A. W. Lievens and P. C. Hoch 1999). It occurs as well along the Mediterranean coast of Africa, through East Africa to southern Africa, and in the Canary and Cape Verde Islands (Raven 1967).
Epilobium hirsutum exceeds almost all other species of the genus in stature, so its size, very large flowers, and densely villous aspect make it easy to identify. R. L. Stuckey (1970) provided a detailed account of the introduction and spread of E. hirsutum in North America, noting the earliest known collection (July 1829) was from Newport, Rhode Island. Most early collections appeared in waste areas, particularly near harbor ballast piles, although some may have been grown in gardens. By the 1890s this species was well established along the Atlantic coastal region from New Jersey and Philadelphia through New England, and around Niagara Falls in the Great Lakes region. During the twentieth century, E. hirsutum spread extensively in southern Ontario and Quebec, south along the Atlantic coast to Maryland, and to all of the states along the southern shores of the Great Lakes, most recently including Wisconsin (1970), and Indiana (1972). It occurs in much the same habitat as that of another, more widely publicized invader, Lythrum salicaria, and sometimes is recorded as a companion species. The earliest known collection in western North America was made in 1933 in Bingen (Klickitat County), Washington. Whether from that introduction or others, E. hirsutum is now naturalized and widespread in the Pacific Northwest. It also was reported recently from the Denver region in Colorado and near Midway in Utah.
Epilobium grandiflorum F. H. Wiggers and E. grandiflorum Allioni are illegitimate names that pertain here.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Epilobium coloratum is highly unusual in having a distribution primarily in southeastern North America. In its overall morphology, particularly leaves and flowers, it more closely resembles European species such as E. obscurum Schreber or E. tetragonum Linnaeus than any other North American species, and shares with the European species the widespread BB chromosome arrangement. There is a general resemblance between E. coloratum and the very widespread E. ciliatum (AA chromosome arrangement); hybrids between these species, known as E. ×wisconsinense Ugent, are highly sterile due to different chromosome arrangements in the parental species. The true affinities of E. coloratum are uncertain. Its occurrence on Hispaniola is also unique and possibly due to a recent introduction.
Prior to 1950, all major treatments of Epilobium attributed the name of E. coloratum to Muhlenberg (as E. coloratum Muhlenberg ex Wildenow), including C. Haussknecht (1884), W. Trelease (1891), and others. However, M. L. Fernald (1945d) noted that Index Kewensis had overlooked the 1807 publication by Biehler and virtually all treatments since that time follow the interpretation by Fernald.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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