The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

codlins-and-cream, fiddle grass, fiddle grass willowherb, great or great hairy willowherb, great willowherb, hairy willlowherb, hairy willow-herb, épilobe hirsute

pygmy willowherb, smooth boisduvalia, smooth spike-primrose, smooth willowherb

Habit Herbs usually robust and rank, sometimes woody near base, with thick, ropelike stolons to 1 m with scattered cataphylls and, often, terminal leafy rosette. Herbs with 1 or more unbranched taproots.
Stems

erect to ascending, often clumped, terete, 25–120(–250) cm, unusually thick, 3–9 mm diam., well branched mainly in distal 1/2, densely long-villous throughout, usually mixed glandular puberulent distally, rarely sparsely villous or densely white-tomentose.

usually suberect, rarely matted, terete, 1.5–50 cm, often with sprawling, decumbent proximal branches, rarely simple, glabrous proximally or throughout, sometimes ± densely strigillose and/or villous distally.

Leaves

opposite proximal to inflorescence, alternate distally, sessile and ± clasping stem;

blade elliptic-lanceolate to narrowly obovate or elliptic, 4–12(–23) × 0.3–4(–5) cm, base cuneate to attenuate, margins serrulate, 15–50 teeth per side, veins 6–9 per side, apex acute to acuminate or obtuse proximally, surfaces ± densely villous;

bracts moderately reduced.

crowded, subsessile, blade lanceolate to narrowly lanceolate or oblong, 0.8–3.5 × 0.2–0.6(–1) cm, longer than subtending internodes, base cuneate, margins evenly serrulate, 4–7 teeth per side, lateral veins obscure, 2–5 per side, apex acute, surfaces strigillose and ± villous, at least along veins and margins;

bracts not much reduced.

Inflorescences

erect racemes or panicles, usually densely villous and glandular puberulent, rarely tomentose.

erect spikes, congested, unbranched, densely strigillose and ± villous or subglabrous.

Flowers

erect;

buds 5–9 × 1.8–4.5 mm, sometimes beaked;

pedicel 3–11 mm;

floral tube 1.3–2.9 × 2.2–4 mm, conspicuous ring of spreading hairs near mouth inside;

sepals oblong-linear, often keeled, 6–12 × 1–3 mm, abaxial surface densely pubescent;

petals bright pink to rose-purple, rarely white, broadly obcordate, 9–20 × 7–15 mm, apical notch 1–3 mm;

filaments white or pink, those of longer stamens 5–10 mm, those of shorter ones 2.5–6 mm;

anthers cream, 1.5–3 × 0.6–1.2 mm;

ovary 15–34 mm, densely villous and glandular puberulent;

style white or pink, 5–12 mm, usually glabrous, stigma deeply 4-lobed, 1.8–2.2 × 3–5.5 mm, lobes recurved or spreading, exserted beyond anthers.

erect, often hidden by subtending bracts, often cleistogamous;

buds 1.2–2 × 0.7–1.1 mm;

floral tube 0.3–1.1 × 0.2–0.8 mm, raised ciliate ring proximal to mouth inside;

sepals reddish green, 0.7–1.9 × 0.6–1.2 mm;

petals pale pink, fading purplish rose, 0.9–3.5 × 0.7–0.9 mm, apical notch 0.3–1.3 mm;

filaments light pink, those of longer stamens 0.5–1.5 mm, those of shorter ones 0.4–0.9 mm;

anthers pale yellow, 0.4–0.8 × 0.3–0.5 mm;

ovary 3–5 mm, usually densely villous;

style pale pink, 0.6–1.8 mm, stigma clavate, irregularly 4-lobed to subentire, 0.5–1 × 0.2–0.6 mm, surrounded by longer anthers.

Capsules

often flushed purple, 25–90 mm, surfaces usually densely villous and glandular puberulent, rarely glabrescent;

pedicel 5–20 mm.

cylindrical to subfusiform, ± terete, 4.5–8 mm, beak 0.8–1 mm, usually dehiscing on distal 1/3, sometimes tardily splitting to base, central axis prematurely disintegrating, villous;

subsessile.

Seeds

narrowly obovoid, 0.8–1.2 × 0.3–0.6 mm, chalazal collar inconspicuous, dark brown, surface coarsely papillose;

coma easily detached, tawny or dull white, 7–10 mm.

7–14 per tightly packed row, irregularly angular-fusiform, 1–1.3 × 0.4–0.6 mm, chalazal collar absent, surface irregularly reticulate.

2n

= 36.

= 30.

Epilobium hirsutum

Epilobium campestre

Phenology Flowering Jun–Sep. Flowering May–Sep.
Habitat Low wet areas along streams, rivers, ponds, and lakes, roadside ditches, along railroad tracks, marshes and swampy areas. Vernally moist flats, depressions, shores, and open fields, usually clay soils.
Elevation 0–150[–3000] m. (0–500[–9800] ft.) 30–3000 m. (100–9800 ft.)
Distribution
from FNA
CO; CT; IL; IN; KY; MA; MD; ME; MI; NH; NJ; NY; OH; OR; PA; RI; UT; VT; WA; WI; WV; BC; NB; NS; ON; PE; QC; Eurasia; Africa [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; ID; MT; ND; NM; NV; OR; SD; UT; WA; WY; AB; BC; SK; Mexico (Baja California); South America (Argentina)
[WildflowerSearch map]
[BONAP county map]
Discussion

Epilobium hirsutum is very widespread in cool temperate Eurasia and montane regions, occurring throughout Europe except in the far north (P. H. Raven 1968), through the Caucasus and central Asia (E. I. Steinberg 1949) to Nepal (Raven 1962), China (Chen C. J. et al. 1992), and Japan (A. W. Lievens and P. C. Hoch 1999). It occurs as well along the Mediterranean coast of Africa, through East Africa to southern Africa, and in the Canary and Cape Verde Islands (Raven 1967).

Epilobium hirsutum exceeds almost all other species of the genus in stature, so its size, very large flowers, and densely villous aspect make it easy to identify. R. L. Stuckey (1970) provided a detailed account of the introduction and spread of E. hirsutum in North America, noting the earliest known collection (July 1829) was from Newport, Rhode Island. Most early collections appeared in waste areas, particularly near harbor ballast piles, although some may have been grown in gardens. By the 1890s this species was well established along the Atlantic coastal region from New Jersey and Philadelphia through New England, and around Niagara Falls in the Great Lakes region. During the twentieth century, E. hirsutum spread extensively in southern Ontario and Quebec, south along the Atlantic coast to Maryland, and to all of the states along the southern shores of the Great Lakes, most recently including Wisconsin (1970), and Indiana (1972). It occurs in much the same habitat as that of another, more widely publicized invader, Lythrum salicaria, and sometimes is recorded as a companion species. The earliest known collection in western North America was made in 1933 in Bingen (Klickitat County), Washington. Whether from that introduction or others, E. hirsutum is now naturalized and widespread in the Pacific Northwest. It also was reported recently from the Denver region in Colorado and near Midway in Utah.

Epilobium grandiflorum F. H. Wiggers and E. grandiflorum Allioni are illegitimate names that pertain here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Epilobium campestre is widespread in temperate western North America. Like E. cleistogamum, it also grows in habitats that are only moist early in the growing season, or otherwise ephemeral moist places, like shores of reservoirs with fluctuating water levels(P. H. Raven and D. M. Moore 1965), and consequently flowers earlier than most species of Epilobium.

The occurrence of this species in Chubut Province, Argentina, appears to be the result of natural long-distance dispersal, probably by birds.

Seeds of Epilobium campestre are inclined about 20° from vertical, which while unique and characteristic is a difficult character to observe. Seeds are tightly packed in rigid capsules, as described under E. cleistogamum.

Oenothera pygmaea Spegazzini 1899, an illegitimate name (not Douglas 1832), pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobium Onagraceae > subfam. Onagroideae > tribe Epilobieae > Epilobium > sect. Epilobiopsis
Sibling taxa
E. anagallidifolium, E. arcticum, E. brachycarpum, E. campestre, E. canum, E. ciliatum, E. clavatum, E. cleistogamum, E. coloratum, E. davuricum, E. densiflorum, E. densum, E. foliosum, E. glaberrimum, E. hallianum, E. hornemannii, E. howellii, E. lactiflorum, E. leptocarpum, E. leptophyllum, E. luteum, E. minutum, E. mirabile, E. montanum, E. nevadense, E. nivium, E. obcordatum, E. obscurum, E. oreganum, E. oregonense, E. pallidum, E. palustre, E. parviflorum, E. rigidum, E. saximontanum, E. septentrionale, E. siskiyouense, E. smithii, E. suffruticosum, E. torreyi
E. anagallidifolium, E. arcticum, E. brachycarpum, E. canum, E. ciliatum, E. clavatum, E. cleistogamum, E. coloratum, E. davuricum, E. densiflorum, E. densum, E. foliosum, E. glaberrimum, E. hallianum, E. hirsutum, E. hornemannii, E. howellii, E. lactiflorum, E. leptocarpum, E. leptophyllum, E. luteum, E. minutum, E. mirabile, E. montanum, E. nevadense, E. nivium, E. obcordatum, E. obscurum, E. oreganum, E. oregonense, E. pallidum, E. palustre, E. parviflorum, E. rigidum, E. saximontanum, E. septentrionale, E. siskiyouense, E. smithii, E. suffruticosum, E. torreyi
Synonyms Chamaenerion hirsutum, E. amplexicaule, E. aquaticum, E. hirsutum var. villosum, E. villosum Boisduvalia campestris, B. glabella, B. glabella var. campestris, B. pygmaea, E. pygmaeum, Oenothera glabella
Name authority Linnaeus: Sp. Pl. 1: 347. (1753) (Jepson) Hoch & W. L. Wagner: Syst. Bot. Monogr. 83: 208. (2007)
Web links