Elymus wiegandii |
Elymus virginicus |
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northern riverbank wildrye, Wiegand's wild-rye, élyme de Wiegand |
common eastern wild-rye, Virginia wild rye, élyme de virginie |
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Habit | Plants cespitose, not rhizomatous, somewhat glaucous. | Plants cespitose, not rhizomatous, sometimes glaucous, especially in the spikes. | ||||||||||||
Culms | 100-180(220) cm, erect; nodes 9-16, mostly concealed by the leaf sheaths, glabrous. |
30-130 cm, erect to slightly decumbent; nodes 4-9, concealed or exposed, usually glabrous, rarely pubescent. |
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Leaves | evenly distributed; sheaths usually glabrous, occasionally villous, often reddish brown; auricles 1-3 mm, brown; ligules to 1 mm; blades (8)10-20(24) mm wide, flat, lax, dark green, adaxial surfaces usually thinly pilose, with weakly spreading hairs on the veins at least near the margins, sometimes villous or glabrous. |
evenly distributed; sheaths usually glabrous, rarely hirsute, occasionally reddish or purplish; auricles absent or to 1.8 mm, pale brown; ligules shorter than 1 mm; blades 2-14(18) mm wide, usually spreading or lax, sometimes becoming involute, basal blades similar to the upper blades, adaxial surfaces usually smooth, sometimes scabridulous, usually glabrous, occasionally pubescent. |
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Spikes | 10-30 cm long, 3-5 cm wide, pendent, the bases often barely exserted, with 2 spikelets per node; internodes 5-8(12) mm long, 0.2-0.3 mm thick at the thinnest sections, usually pubescent beneath the spikelets. |
(3)4-16(22) cm long, 1-2.2(2.5) cm wide, erect, the bases often sheathed, with 2 spikelets per node, rarely with 3 at some nodes; internodes 3-5 mm long, 0.25-0.5 thick at the thinnest sections, smooth and glabrous, or scabrous, or with hairs beneath the spikelets. |
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Spikelets | 12-20 mm, divergent, with (3)4-6(7) florets, lowest florets functional; disarticulation above the glumes and beneath each floret. |
10-15 mm, appressed to slightly divergent, with (2)3-4(6) florets, lowest florets functional; disarticulation below the glumes and each floret, or the lowest floret falling with the glumes. |
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Glumes | equal or subequal, 12-30 mm including the often undifferentiated awns, the basal 0.5-1 mm subterete and slightly indurate, glume bodies 7-12 mm long, (0.2)0.4-0.9(1.1) mm wide, linear-setiform, entire, widening or parallel-sided above the base, 1-3(5)-veined, glabrous, hispidulous or villous, especially near the margins, margins firm, awns (5)8-15(18) mm, straight or flexuous; lemmas 10-15 mm, usually uniformly appressed-villous, rarely scabrous-hirtellous or glabrous, awns 15-25(30) mm, moderately to strongly outcurving; paleas 9-14 mm, narrowly truncate, minutely bidentate; anthers 2-3.5 mm. |
subequal or equal, the basal 1-4 mm terete, indurate, without evident venation, bowed out, yellowish, glume bodies 7-15 mm long, (0.5)0.7-2.3 mm wide, linear-lanceolate, widening above the base, 3-5(8)-veined, usually smooth or scabridulous, margins firm, awns 3-10(15) mm, straight; lemmas 6-10 mm, scabridulous, glabrous or villous-hirsute, awns (5)8-20(25) mm, straight; paleas 5-9 mm, obtuse; anthers 2-3.5(4) mm. |
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Anthesis | from mid-July to early August. |
usually mid-June to late July (mid-August). |
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2n | = 28. |
= 28. |
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Elymus wiegandii |
Elymus virginicus |
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Distribution |
CT; IA; KY; MA; ME; MI; MN; ND; NH; NJ; NY; PA; SD; VT; WI; WY; NB; ON; QC; SK
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AL; AR; AZ; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; AB; BC; LB; MB; NB; NS; ON; PE; QC; SK
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Discussion | Elymus wiegandii grows in moist or damp, rich, alluvial soil, especially on sandy river terraces and in woods and thickets, primarily from Saskatchewan through much of the Great Lakes region to Nova Scotia and Connecticut. It has abnormal neocentric chromosomes with meiotic irregularities that appear to limit the fertility of its hybrids, and even some crosses within the species (Vilkomerson 1950). It may be derived from hybrids between E. canadensis (p. 303) and perhaps E. riparius (p. 302). The latter species is similar to E. wiegandii and overlaps with it in range and habitat within the Great Lakes region, where there are a few plants that appear to be hybrids between the two. Plants with scabrous-hirtellous or glabrous lemmas (E. wiegandii f. calvescens Fernald) are known from Maine and New Hampshire. Elymus wiegandii is often confused with sympatric E. canadensis and E. diversiglumis (p. 316), but it has a distinctive robust, broad-leaved habit. It is intermediate between the two in spike density and glume development. Occasional plants with glabrous leaves and less pendent spikes suggest introgression from E. canadensis, but artificial crosses produced no fertile Fj plants (Church 1958). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Elymus virginicus is widespread in temperate North America, growing as far west as British Columbia and Arizona. It is infrequent to rare in the Rocky Mountains, western Great Plains, and southeastern coastal plain. It is a complex species, divided here into four intergrading varieties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 305. | FNA vol. 24, p. 298. | ||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | ||||||||||||
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Synonyms | E. canadensis var. wiegandii | |||||||||||||
Name authority | Fernald | L. | ||||||||||||
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