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northern riverbank wildrye, Wiegand's wild-rye, élyme de Wiegand

wildrye

Habit Plants cespitose, not rhizomatous, somewhat glaucous. Plants loosely cespitose, without conspicuous rhizomes.
Culms

100-180(220) cm, erect;

nodes 9-16, mostly concealed by the leaf sheaths, glabrous.

30-130 cm tall, 1-5 mm thick, erect or weakly decumbent;

nodes 3-4, glabrous, glaucous.

Leaves

evenly distributed;

sheaths usually glabrous, occasionally villous, often reddish brown;

auricles 1-3 mm, brown;

ligules to 1 mm;

blades (8)10-20(24) mm wide, flat, lax, dark green, adaxial surfaces usually thinly pilose, with weakly spreading hairs on the veins at least near the margins, sometimes villous or glabrous.

basal and cauline;

sheaths glaucous, glabrous or with hairs, lower sheaths sometimes hairy, upper sheaths glabrous, margins sometimes ciliate;

auricles 1.5-2.5 mm;

ligules about 0.3 mm;

blades 10-25 cm long, 3-10 mm wide, glabrous or pilose.

Spikes

10-30 cm long, 3-5 cm wide, pendent, the bases often barely exserted, with 2 spikelets per node;

internodes 5-8(12) mm long, 0.2-0.3 mm thick at the thinnest sections, usually pubescent beneath the spikelets.

10-22 cm long, 1.5-2.5 cm wide including the awns, 0.8-1 cm wide excluding the awns, inclined to nodding, with 1 spikelet at all or most nodes;

rachises scabrous on the edges, glabrous below the spikelets;

internodes 10-25 mm.

Spikelets

12-20 mm, divergent, with (3)4-6(7) florets, lowest florets functional;

disarticulation above the glumes and beneath each floret.

5-22 mm long, about 5 mm wide, appressed, with 4-12 florets;

disarticulation above the glumes, beneath the florets.

Glumes

equal or subequal, 12-30 mm including the often undifferentiated awns, the basal 0.5-1 mm subterete and slightly indurate, glume bodies 7-12 mm long, (0.2)0.4-0.9(1.1) mm wide, linear-setiform, entire, widening or parallel-sided above the base, 1-3(5)-veined, glabrous, hispidulous or villous, especially near the margins, margins firm, awns (5)8-15(18) mm, straight or flexuous;

lemmas 10-15 mm, usually uniformly appressed-villous, rarely scabrous-hirtellous or glabrous, awns 15-25(30) mm, moderately to strongly outcurving;

paleas 9-14 mm, narrowly truncate, minutely bidentate;

anthers 2-3.5 mm.

narrowly elliptic to lance-oblong, apices acute to acuminate;

lower glumes 5-11 mm;

upper glumes 7-13 mm;

lemmas 7-12 mm, mostly glabrous, glabrate, or sparsely hairy, margins with coarse stiff hairs, hairs to 1 mm, apices abruptly narrowed, awned, awns 10-20 mm, scabrous, strongly outcurved to recurved;

paleas 2/3 - 4/5 the length of the lemmas, keels winged distally, distinctly outwardly curved below the apices, apices 0.5-0.6 mm wide, truncate to rounded;

anthers about 2 mm.

Anthesis

from mid-July to early August.

Genome

StY.

2n

= 28.

= 28.

Elymus wiegandii

Elymus ciliaris

Distribution
from FNA
CT; IA; KY; MA; ME; MI; MN; ND; NH; NJ; NY; PA; SD; VT; WI; WY; NB; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
[BONAP county map]
Discussion

Elymus wiegandii grows in moist or damp, rich, alluvial soil, especially on sandy river terraces and in woods and thickets, primarily from Saskatchewan through much of the Great Lakes region to Nova Scotia and Connecticut. It has abnormal neocentric chromosomes with meiotic irregularities that appear to limit the fertility of its hybrids, and even some crosses within the species (Vilkomerson 1950). It may be derived from hybrids between E. canadensis (p. 303) and perhaps E. riparius (p. 302). The latter species is similar to E. wiegandii and overlaps with it in range and habitat within the Great Lakes region, where there are a few plants that appear to be hybrids between the two. Plants with scabrous-hirtellous or glabrous lemmas (E. wiegandii f. calvescens Fernald) are known from Maine and New Hampshire.

Elymus wiegandii is often confused with sympatric E. canadensis and E. diversiglumis (p. 316), but it has a distinctive robust, broad-leaved habit. It is intermediate between the two in spike density and glume development. Occasional plants with glabrous leaves and less pendent spikes suggest introgression from E. canadensis, but artificial crosses produced no fertile Fj plants (Church 1958).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus ciliaris is native to northern China and Japan. It was collected from ballast dumps in Portland, Oregon, in 1899 and 1902; it is not established in the Flora region. A.S. Hitchcock identified both specimens on the sheet (US 1017954) as Agropyron caninum (L.) P. Beauv. [= Elymus caninus, p. 322], from which E. ciliaris differs in its short, rounded paleas and relatively short glumes with distinctly outwardly curving keels. The other specimen on that sheet is E. tsuskushiensis (p. 336).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 305. FNA vol. 24, p. 336.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Synonyms E. canadensis var. wiegandii
Name authority Fernald (Trin.) Tzvelev
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