Elymus stebbinsii
Elymus hystrix
Parish wheatgrass, Stebbins' wheat grass, Stebbins' wildrye
bottlebrush grass, eastern bottle-brush grass, glumeless wlldrye
60-140 cm;
nodes glabrous or retrorsely pubescent.
50-140 cm, usually erect, occasionally geniculate below;
nodes 4-8, exposed or concealed, glabrous.
evenly distributed;
sheaths glabrous or pubescent;
auricles usually present, 0.5-2 mm;
ligules 0.3-3.5 mm, truncate to acute, sometimes long-ciliate;
blades 4-6.5 mm wide, flat or the margins involute, straight.
evenly distributed;
sheaths usually glabrous, occasionally pilose, often purplish;
auricles usually present, 0.5-3 mm, brown to black;
ligules 1-2(3) mm;
blades 4-16 mm wide, lax, usually deep glossy green, adaxial surfaces pilose or scabridulous.
15-31 cm long, 0.4-1.5 cm wide including the awns, 0.4-0.8 cm wide excluding the awns, erect, with 1 spikelet per node;
internodes 9-27 mm long, 1-1.3 mm wide, glabrous, smooth.
7-20 cm long, 4-7 cm wide, more or less erect, usually with 2 spikelets per node, rarely with 3 at some nodes;
internodes (3)4-8(10) mm long, (0.1)0.2-0.3(0.4) mm thick at the thinnest sections, flexuous, usually glabrous, sometimes scabrous or hirsute, usually with green lateral bands.
13-29 mm long, from shorter than to almost twice as long as the internodes, 2.5-5 mm wide, appressed, with 5-7 florets;
rachillas glabrous;
disarticulation above the glumes and beneath each floret.
10-18 mm, strongly divergent to patent at maturity, with (1)2-4(6) florets, lowest florets functional;
disarticulation above the glumes, beneath each floret.
subequal, 7.5-12 mm long, 1.2-1.5 mm wide, lanceolate, widest at about mid-length, flat or rounded on the back, 5-veined, veins smooth, scabrous or just the midvein scabridulous, margins widest at about midlength, apices acute, unawned;
lemmas 9-12 mm, glabrous, sometimes scabrous, acute, unawned or awned, awns to 28 mm, straight;
paleas subequal to the lemmas, tapering, apices 0.2-0.3 mm wide;
anthers (3.5)4-7 mm.
usually vestigial, sometimes 1-3 mm long, about 0.1 mm wide, subulate, entire, with no evident veins, occasionally to 10(20) mm long including the undifferentiated awns and differing in length by more than 5 mm, 0.1-0.2 mm wide, setaceous, tapering from the base, usually glabrous, occasionally appressed-puberulent to strigose, sometimes scabrous, usually straight, rarely somewhat curving, margins firm;
lemmas 8-11 mm, usually glabrous, occasionally appressed-puberulent to strigose, especially near the margins and apices, awns (12)20-40(47) mm, usually straight, rarely somewhat curving;
paleas 7-11 mm, obtuse or truncate, occasionally emarginate;
anthers 2.5-5 mm.
mid-June to early July.
= 28.
= 28.
Elymus stebbinsii
Elymus hystrix
Elymus stebbinsii is restricted to California, where it grows on dry slopes, chaparral, and wooded areas, at elevations below 1600 m. It differs from other Elymus species primarily in its combination of long anthers and solitary spikelets. It is often confused with E. glaucus (p. 306) and E. trachycaulus (p. 321) with solitary spikelets. It differs from both in its longer anthers, and from most representatives of E. glaucus in its acute, but unawned, glumes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Elymus hystrix grows in dry to moist soils in open woods and thickets, especially on base-rich slopes and small stream terraces. It grows throughout most of temperate eastern North America, extending west to Manitoba and Oklahoma, but is absent from the southern portion of the coastal plain.
Plants with pubescent lemmas have been recognized as Elymus hystrix var. bigelovianus (Fernald) Bowden. These occur infrequently north of a line from South Dakota through Kentucky to New Jersey, and are often mixed with the typical variety; uniform populations are known in the northeastern United States. Plants with pubescent blades are also more prevalent to the north. Elymus hystrix hybridizes with most eastern species of Elymus. Introgression may account for the considerable variation in glume development and spikelet appression among these species. Lack of glumes may be a recessive character, with even slight glume development indicating introgression (Church 1967b). Plants with relatively well-developed, subequal glumes are presumed to be of hybrid origin. Such plants include most material from the Carolina piedmont region, where E. glabriflorus (p. 296) is the most likely source of introgression. The relatively frequent hybrids with E. virginicus (p. 298) are usually sterile, but Church (1967b) made crosses through three segregating generations. Within the ranges of E. diversiglumis (p. 316), E. svensonii (p. 314), and E. churchii (p. 314;, there appear to be frequent introgressants between these species and E. hystrix. Further east, especially in the Appalachian regions of North Carolina, Virginia, West Virginia, and Maryland (including the shale barrens and nearby), there are scattered plants of E. hystrix with curving awns and, in a few cases, appressed spikelets (Campbell 2002). Whether these represent occasional variation within the E. hystrix gene pool, or whether they are outlying remnants of introgression with E. canadensis (p. 303) during a past eastward extension, is unknown.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Lemmas awned, awns 8-28 mm long; lower leaf sheaths rarely pubescent; spikelets 13-22 mm long | subsp. septentrionalis |
1. Lemmas unawned or with awns to 8(12) mm long; lower leaf sheaths pubescent or glabrous; spikelets 17-29 mm long | subsp. stebbinsii |
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