Elymus scribneri |
Elymus tsukushiensis |
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Scribner's wheat grass, Scribner's wild rye, spreading wheatgrass |
tsukushi wildrye |
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Habit | Plants cespitose, not rhizomatous. | Plants loosely cespitose, without conspicuous rhizomes. |
Culms | 15-35(55) cm, prostrate to strongly decumbent, at least at the base; nodes glabrous. |
25-100 cm tall, 1.3-3.5 mm thick, erect; nodes 4-6, glabrous. |
Sheaths | glabrous or shortly pilose; auricles usually present, 0.5-1 mm; ligules 0.2-0.4(0.7) mm, usually truncate, occasionally acute, entire to erose; blades 1.5-4 mm wide, usually involute, adaxial surfaces prominently ribbed. |
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Leaves | basal and cauline; sheaths glaucous, glabrous or with hairs, margins glabrous or ciliate distally; auricles 1-2 mm; ligules 0.2-0.7 mm, truncate; blades 3-10 mm wide, flattish, often glaucous. |
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Spikes | 3.5-10 cm long, 0.8-1.2 cm wide excluding the awns, 3-6 cm wide including the awns, usually with 1 spikelet per node, occasionally with 2 spikelets at the lower nodes; internodes 2.5-5(7) mm long, 0.5-1 mm wide, glabrous, mostly smooth, edges scabrous. |
(6.5)10-25 cm long, 1.4-4 cm wide including the awns, 0.7-20 cm wide excluding the awns, flexuous, nodding; rachises densely to sparsely hirsute on the edges, hairs about 0.2 mm, glabrous elsewhere, glaucous; internodes (5)8-20 mm. |
Spikelets | 9-15 mm long, 6-12 mm wide, appressed to ascending, with 3-6 florets; rachilla internodes 0.8-1.3 mm, scabridulous; disarticulation initially at the rachis nodes, subsequently beneath each floret. |
15-25 mm, loosely appressed or ascending, with 5-10 florets; rachillas hairy, hairs about 0.1 mm; disarticulation above the glumes, beneath the florets. |
Glumes | 4-9 mm long, 0.5-1 mm wide, mostly glabrous, midveins scabrous, 3-5-veined, entire, tapering into a divergent, 12-30 mm awn; lemmas 7-10 mm, usually glabrous, occasionally scabridulous, awned, awns 15-30 mm, divergent, scabridulous; paleas usually longer than the lemmas, apices ciliate, truncate or the veins extending into teeth, teeth about 0.5 mm; anthers 1-1.6 mm. |
lanceolate, tapering from about midlength, adaxial surfaces glabrous, hyaline margins about 0.1 mm wide, strongly keeled distally, midvein scabrous distally, other veins smooth or scabrous, apices acute to acuminate, sometimes awned, awns 2-5 mm; lower glumes 4-7 mm, 3-5-veined; upper glumes 5-8 mm, 5-veined; calluses glabrous; lemmas 8-12 mm, lanceolate, glabrous or pilose, apices acute, awned, awns 20-40 mm, straight or flexuous; paleas from slightly shorter than to longer than the lemmas, keels narrowly winged distally, not or scarcely extending beyond the intercostal region, distinctly outwardly curved below the apices, apices 0.3-0.5 mm wide; anthers 1.5-2.5 mm. |
2n | = 28. |
= 42. |
Elymus scribneri |
Elymus tsukushiensis |
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Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB
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Discussion | Elymus scribneri grows in rocky areas in open subalpine and alpine regions, at 2500-3200 m, often in windswept locations, in southwestern Alberta and the western United States. It is often confused with E. elymoides (p. 318), but differs from that species in having only one spikelet per node, wider glumes, and more tardily disarticulating rachises. It also resembles E. sierrae (see next), from which it differs in its disarticulating rachises, denser spikes, and shorter anthers. Dewey (1963) concluded that Elymus trachycaulus subsp. andinus consists of hybrids between E. scribneri and E. trachycaulus (p. 321). In addition, several taxonomists have suggested that E. scribneri consists of fertile hybrids between E. violaceus (p. 324) and E. elymoides. This suggestion is supported by the frequency with which the three taxa are sympatric, the morphological variation exhibited by E. scribneri, and cytogenetic data (Dewey 1967). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Elymus tsuskushiensis is native to northeastern China, Japan, and Korea. It was collected from ballast dumps in Portland, Oregon, but is not established in the Flora region. Hitchcock (1951) identified it and E. ciliaris as Agropyron caninum (L.) P. Beauv. [= Elymus caninus, p. 322], but that species has flatter glumes that are longer in relation to the lemmas than those of E. tsuskushiensis, and paleas with straight or slightly outwardly curved keels. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 24, p. 330. | FNA vol. 24, p. 336. |
Parent taxa | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus | Poaceae > subfam. Pooideae > tribe Triticeae > Elymus |
Sibling taxa | ||
Synonyms | Agropyron scribneri | |
Name authority | (Vasey) M. E. Jones | Honda |
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