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Scribner's wheat grass, Scribner's wild rye, spreading wheatgrass

northern wheatgrass, tuft wheatgrass

Habit Plants cespitose, not rhizomatous. Plants cespitose, sometimes appearing weakly rhizomatous.
Culms

15-35(55) cm, prostrate to strongly decumbent, at least at the base;

nodes glabrous.

35-100 cm, ascending to erect;

nodes sometimes pubescent.

Sheaths

glabrous or shortly pilose;

auricles usually present, 0.5-1 mm;

ligules 0.2-0.4(0.7) mm, usually truncate, occasionally acute, entire to erose;

blades 1.5-4 mm wide, usually involute, adaxial surfaces prominently ribbed.

glabrous;

auricles absent;

ligules 0.5-1 mm, truncate to rounded;

blades 3-10 mm wide, flat, usually glabrous, abaxial surfaces smooth or scabridulous, adaxial surfaces scabrous.

Spikes

3.5-10 cm long, 0.8-1.2 cm wide excluding the awns, 3-6 cm wide including the awns, usually with 1 spikelet per node, occasionally with 2 spikelets at the lower nodes;

internodes 2.5-5(7) mm long, 0.5-1 mm wide, glabrous, mostly smooth, edges scabrous.

5-20 cm long, 0.4-0.8 cm wide, erect, with 1 spikelet per node;

internodes 7-8 mm long, about 0.5 mm wide, glabrous below the spikelets.

Spikelets

9-15 mm long, 6-12 mm wide, appressed to ascending, with 3-6 florets;

rachilla internodes 0.8-1.3 mm, scabridulous;

disarticulation initially at the rachis nodes, subsequently beneath each floret.

12-20 mm, appressed, with 4-7 florets;

rachillas hairy, hairs 0.3-0.5 mm;

disarticulation above the glumes, beneath each floret.

Glumes

4-9 mm long, 0.5-1 mm wide, mostly glabrous, midveins scabrous, 3-5-veined, entire, tapering into a divergent, 12-30 mm awn;

lemmas 7-10 mm, usually glabrous, occasionally scabridulous, awned, awns 15-30 mm, divergent, scabridulous;

paleas usually longer than the lemmas, apices ciliate, truncate or the veins extending into teeth, teeth about 0.5 mm;

anthers 1-1.6 mm.

6-10 mm long, 1/3-2/3 the length of the spikelets and to about 1/2 the length of the adjacent lemmas, 0.8-1.8 mm wide, widest at about midlength, lanceolate, flat, rounded, or symmetrically keeled, usually green or green tinged with purple, 3-4-veined, veins scabridulous, scabrous, or with hairs to 0.3 mm, usually glabrous elsewhere, margins subequal, about 0.3 mm wide, widest near midlength, apices acute, unawned or awned, awns to 1 mm;

lemmas 8-12 mm, hairy throughout or glabrous distally, hairs all alike, 0.2-0.3 mm, apices unawned or awned, awns to 7 mm, straight;

paleas subequal to the lemmas, tapering to the apices, apices about 0.8 mm wide;

anthers 1-2 mm.

2n

= 28.

= 28.

Elymus scribneri

Elymus macrourus

Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; YT
[BONAP county map]
Discussion

Elymus scribneri grows in rocky areas in open subalpine and alpine regions, at 2500-3200 m, often in windswept locations, in southwestern Alberta and the western United States. It is often confused with E. elymoides (p. 318), but differs from that species in having only one spikelet per node, wider glumes, and more tardily disarticulating rachises. It also resembles E. sierrae (see next), from which it differs in its disarticulating rachises, denser spikes, and shorter anthers.

Dewey (1963) concluded that Elymus trachycaulus subsp. andinus consists of hybrids between E. scribneri and E. trachycaulus (p. 321). In addition, several taxonomists have suggested that E. scribneri consists of fertile hybrids between E. violaceus (p. 324) and E. elymoides. This suggestion is supported by the frequency with which the three taxa are sympatric, the morphological variation exhibited by E. scribneri, and cytogenetic data (Dewey 1967).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus macrourus grows on river banks and bars, lake shores, and hillsides in northwestern North America. Outside of North America, it grows across the Russian arctic, and extends south into the boreal forest. Plants growing on shifting river banks and bars often appear rhizomatous, as the lower internodes elongate in response to the disturbed substrate. Plants of E. macrourus differ from E. alaskanus (see next) in the shape of their glumes and their narrower glume margins, and from E. trachycaulus (p. 321) in their relatively short glumes and evidently hairy rachilla segments.

Three varieties of Elymus macrourus are recognized in Russian treatments. It is not clear to which, if any, of the Russian varieties North American plants belong. A circumboreal study is needed, using plants grown from seeds collected in the wild. Seeds available as E. macrourus through germplasm resources appear to be misidentified.

Elymus macrourus is one of the parents in both E. ×palmerensis (p. 340) and ×Elyhordeum pilosilemma (p. 284).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 330. FNA vol. 24, p. 324.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Synonyms Agropyron scribneri Agropyron macrourum, Roegneria turuchanensis, Roegneria macroura, Agropyron sericeum
Name authority (Vasey) M. E. Jones (Turcz. ex Steud.)
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