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Scribner's wheat grass, Scribner's wild rye, spreading wheatgrass

Church's wildrye

Habit Plants cespitose, not rhizomatous. Plants cespitose, not rhizomatous, often somewhat glaucous.
Culms

15-35(55) cm, prostrate to strongly decumbent, at least at the base;

nodes glabrous.

50-120 cm, erect;

nodes 4-8, exposed or concealed, often reddish brown or blackish, glabrous.

Sheaths

glabrous or shortly pilose;

auricles usually present, 0.5-1 mm;

ligules 0.2-0.4(0.7) mm, usually truncate, occasionally acute, entire to erose;

blades 1.5-4 mm wide, usually involute, adaxial surfaces prominently ribbed.

Leaves

evenly distributed;

sheaths usually glabrous, sometimes pubescent at the summit;

auricles 1-2 mm, often reddish brown or blackish;

ligules to 1 mm, often reddish brown;

blades 3-11 mm wide, lax, adaxial surfaces glabrous or short-pilose.

Spikes

3.5-10 cm long, 0.8-1.2 cm wide excluding the awns, 3-6 cm wide including the awns, usually with 1 spikelet per node, occasionally with 2 spikelets at the lower nodes;

internodes 2.5-5(7) mm long, 0.5-1 mm wide, glabrous, mostly smooth, edges scabrous.

10-18 cm long, 3-5 cm wide, slightly nodding, with 2 spikelets per node;

internodes (5)7-13(18) mm long, about 0.2 mm thick at the thinnest sections, flexuous, with green lateral bands, glabrous except the dorsal angles hispid.

Spikelets

9-15 mm long, 6-12 mm wide, appressed to ascending, with 3-6 florets;

rachilla internodes 0.8-1.3 mm, scabridulous;

disarticulation initially at the rachis nodes, subsequently beneath each floret.

10-15 mm, usually appressed, with 3(5) florets, lowest florets functional;

disarticulation above the glumes, beneath each floret.

Glumes

4-9 mm long, 0.5-1 mm wide, mostly glabrous, midveins scabrous, 3-5-veined, entire, tapering into a divergent, 12-30 mm awn;

lemmas 7-10 mm, usually glabrous, occasionally scabridulous, awned, awns 15-30 mm, divergent, scabridulous;

paleas usually longer than the lemmas, apices ciliate, truncate or the veins extending into teeth, teeth about 0.5 mm;

anthers 1-1.6 mm.

often differing in length by more than 5 mm, sometimes vestigial to absent from the upper spikelets or throughout, 0-15(20) mm long including the undifferentiated awns, indurate at the base, 0.1-0.3 mm wide, setaceous to subulate, entire, 0-1-veined, glabrous, margins firm, awns often outcurving;

lemmas 8-10 mm, usually hairy, occasionally glabrous, awns (10)20-30(35) mm, slightly to strongly outcurving at maturity;

paleas 7-9 mm, obtuse to truncate, sometimes emarginate;

anthers 2.5-3 mm, evident in June.

2n

= 28.

= unknown.

Elymus scribneri

Elymus churchii

Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; MO; OK
[BONAP county map]
Discussion

Elymus scribneri grows in rocky areas in open subalpine and alpine regions, at 2500-3200 m, often in windswept locations, in southwestern Alberta and the western United States. It is often confused with E. elymoides (p. 318), but differs from that species in having only one spikelet per node, wider glumes, and more tardily disarticulating rachises. It also resembles E. sierrae (see next), from which it differs in its disarticulating rachises, denser spikes, and shorter anthers.

Dewey (1963) concluded that Elymus trachycaulus subsp. andinus consists of hybrids between E. scribneri and E. trachycaulus (p. 321). In addition, several taxonomists have suggested that E. scribneri consists of fertile hybrids between E. violaceus (p. 324) and E. elymoides. This suggestion is supported by the frequency with which the three taxa are sympatric, the morphological variation exhibited by E. scribneri, and cytogenetic data (Dewey 1967).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus churchii grows in dry, rocky, often relatively base-rich soils, in open woods on ridges, and on bluffs and river banks. Its range includes the central Ouachita Mountains and the western Ozark Mountains in Arkansas, Oklahoma, and Missouri.

Elymus churchii used to be included in E. interruptus (p. 306) (Steyermark 1963; Smith 1991). It is similar to the more eastern, disjunct E. svensonii (see previous), from which it differs in its more open spikes, longer awns, fewer florets per spikelet, and less pubescent, less glaucous foliage. Like E. svensonii, E. churchii may have originated from hybridization between E. canadensis (p. 303) and E. hystrix (p. 316); occasional intermediates with both species exist (Campbell 2002).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 330. FNA vol. 24, p. 314.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Synonyms Agropyron scribneri
Name authority (Vasey) M. E. Jones J.J.N. Campb.
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