Elymus riparius
Elymus ×hansenii
eastern riverbank wild-rye, river wild-rye, riverbank wildrye, élyme des rivages
Hansen squirreltail, Hansen's wheatgrass, Hansen's wildrye
70-160 cm, erect, sometimes rooting at the lower nodes;
nodes 5-10, mostly concealed, glabrous.
60-120 cm.
evenly distributed;
sheaths usually glabrous or scabridulous, often reddish brown;
auricles absent or to 2 mm, brown;
ligules shorter than 1 mm;
blades (5)8-15(25) mm wide, flat, lax, dull green, drying to grayish, adaxial surfaces glabrous or scabrous.
evenly distributed;
sheaths smooth;
ligules to 1 mm;
blades 10-30 cm long, 2-8 mm wide, flat or the margins involute.
7-25 cm long, 2-4 cm wide, nodding, exserted, usually with 2 spikelets per node, rarely with 3 at some nodes;
internodes 3-5(8) mm long, 0.2-0.35 thick at the thinnest sections, usually glabrous below the spikelets.
5-20 cm, straight or nodding, with 2+ spikelets per node;
internodes about 10 mm;
disarticulation in the rachises.
10-20 mm, strongly divergent, with 2-3(4) florets, lowest florets functional;
disarticulation above the glumes and beneath each floret.
about 15 mm, with 3-5 florets.
equal or subequal, 14-30 mm including the sometimes undifferentiated awn, the basal 0.5-2 mm terete, indurate, straight or nearly so, veins not evident, glume bodies 9-17 mm long, (0.3)0.5-0.8(1) mm wide, linear-setiform, entire, widening or parallel-sided above the base, 2-3(4)-veined, usually hispidulous or scabrous, rarely glabrous, margins firm, awns (5)8-18 mm, straight;
lemmas 7-14 mm, usually hispidulous, sometimes scabrous, awns 15-35 mm, usually straight, those of the basal spikelets occasionally contorted;
paleas 6-9 mm, usually acute, sometimes obtuse to truncate, bidentate;
anthers 2-2.7 mm.
narrowly lanceolate, 2-3-veined, awned, awns 25-35 mm;
lemmas 10-12 mm, awned, awns 40-50 mm, outcurving;
paleas subequal to the lemmas, truncate or bidentate.
late June to late July.
= 28.
Elymus riparius
Elymus ×hansenii
Elymus riparius grows in moist, usually alluvial and often sandy soils in woods and thickets, usually along larger streams and occasionally along upland ditches. It is widespread in most of temperate east-central North America. It is rare in southern Ontario and Quebec, and the eastern Great Plains. It is virtually absent from the southeastern coastal plain.
Elymus riparius is relatively uniform and distinct. It is sometimes confused with E. canadensis (see next), but that species has curving awns. It hybridizes occasionally with several other taxa, especially E. virginicus var. virginicus (p. 300) and E. hystrix (p. 316), but the hybrids produce only late, depauperate spikes or none at all (e.g., Church 1958).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Elymus ×hansenii refers to hybrids between E. glaucus (p. 306) and either E. elymoides or E. multisetus (p. 318). It is not clear which of the latter two species is involved. It is a fairly common hybrid in those parts of western North America where both parents grow. The glumes of the type specimen are as wide as those in E. glaucus, and some are divided longitudinally, as in E. elymoides and E. multisetus. As in other hybrids involving E. elymoides and E. multisetus, the rachis of E. ×hansenii disarticulates at maturity.
Elymus ×hansenii is an Elymus named hybrid
Elymus is notorious for its ability to hybridize. Most of its interspecific hybrids are partially fertile, permitting introgression between the parents. The descriptions provided below are restricted to the named interspecific hybrids. They should be treated with caution and some skepticism; some are based solely on the type specimen, because little other reliably identified material was available. Moreover, as the descriptions of the non-hybrid species indicate, many other interspecific hybrids exist.
The parentage of all hybrids is best determined in the field. Perennial hybrids, such as those in Elymus, can persist in an area after one or both parents have died out, but the simplest assumption is that both are present. Interspecific hybrids of Elymus that have disarticulating rachises presumably have E. elymoides or E. multisetus as one of their parents.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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