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big squirreltail, big squirreltail grass

chiendent, chiendent rampant, couch grass, creeping wild-rye, quack grass, quackgrass wildrye

Habit Plants cespitose, not rhizomatous. Plants strongly rhizomatous, sometimes glaucous.
Culms

15-65 cm, erect to ascending, usually puberulent;

nodes 4-6, mostly concealed, glabrous.

50-100 cm.

Leaves

evenly distributed;

sheaths glabrous or white-villous;

auricles usually present, 0.5-1.5 mm;

ligules to 1 mm, truncate, entire or lacerate;

blades 1.5-4(5) mm wide, often ascending and involute, adaxial surfaces scabrous, pilose, or villous.

sometimes somewhat basally concentrated;

sheaths pilose or glabrous proximally;

auricles 0.3-1 mm;

ligules 0.25-1.5 mm;

blades 6-10 mm wide, usually flat, abaxial surfaces glabrous or sparsely pilose, adaxial surfaces usually sparsely pilose over the veins, sometimes glabrous, veins smooth, widely spaced, primary veins prominent, separated by the secondary veins.

Spikes

5-20 cm long, 5-15 cm wide, erect, sometimes partially enclosed at the base, with 2 spikelets per node, rarely with 3-4 at some nodes;

internodes 3-5(8) mm long, 0.1-0.3 mm thick at the thinnest sections, glabrous beneath the spikelets.

5-15 cm long, 0.5-1.5 cm wide, erect, usually with 1 spikelet per node, occasionally with 2 at a few nodes;

internodes 4-6(9.5) mm long, 0.5-1.2 mm wide, smooth or scabrous, glabrous, evenly puberulent, or sparsely pilose, hairs to 0.3 mm.

Spikelets

10-15 mm, divergent, with 2-4 florets, lowest florets sterile and glumelike in 1 or both spikelets at each node;

disarticulation initially at the rachis nodes, subsequently beneath each floret.

10-27 mm, appressed to ascending, with 4-7 florets;

disarticulation above the glumes, beneath each floret.

Glumes

subequal, (10)30-100 mm including the awns, the bases indurate and glabrous, glume bodies (2)5-10 mm long, 1-2 mm wide, setaceous, 2-3-veined, margins firm, awns (8)25-90 mm, each split into 3-9 unequal divisions, scabrous, flexuous to outcurving from near the glume bases at maturity;

fertile lemmas 8-10 mm, smooth or scabrous near the apices, 2 lateral veins extending into bristles to 10 mm, awns (10)20-110 mm long, about 0.2 mm wide at the base, divergent to arcuate;

paleas 7-9 mm, veins usually extending into about 1 mm bristles, apices acute to truncate;

anthers 1-2 mm.

oblong, glabrous, keeled distally, keels inconspicuous and smooth proximally, scabrous and conspicuous distally, lateral veins inconspicuous, hyaline margins present in the distal 1/2, apices acute, unawned or awned, awns to 3 mm;

lower glumes 8.8-11.4 mm, 3-6-veined;

upper glumes 7-12 mm, 5-7-veined;

lemmas 8-12 mm, glabrous, mostly smooth, sometimes scabridulous distally, unawned or with a 0.2-4(10) mm awn, awns straight;

paleas 7-9.5 mm, keels ciliate from 1/2 to almost the entire length, apices emarginate, truncate, or rounded;

anthers 4-7 mm.

Anthesis

from late May to June.

2n

= 28.

= 22, 42.

Elymus multisetus

Elymus repens

Distribution
from FNA
AZ; CA; CO; ID; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Elymus multisetus grows in dry, often rocky, open woods and thickets on slopes and plains, from central Washington and Idaho to southern California, Colorado, and northwestern Arizona, and from sea level to 2000 m. It has also been reported from Baja California, Mexico. It usually grows in less arid habitats than E. elymoides subsp. elymoides (p. 319), but the two taxa are sometimes sympatric.

Wilson (1963) reported a wide belt of introgression between Elymus multisetus and E. elymoides subsp. elymoides from southeastern California to southern Nevada, but not in other areas where they are sympatric. There are also probable hybrids with E. glaucus (p. 306) and Pseudoroegneria spicata (p. 281).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus repens is native to Eurasia; it is now established through much of the Flora region, extending from Alaska to Greenland and south to California, Texas, and North Carolina. It grows well in disturbed sites, spreading rapidly via its long rhizomes, as well as by seed. It is also drought tolerant. Although it is listed a noxious weed in several states, it provides good forage. It differs from E. hoffmannii (see next) in having widely spaced, unequally prominent leaf veins and, usually, shorter awns.

Godley (1947) demonstrated that lemma awn development, glaucousness, and the pubescence of the rachises are each effectively controlled by single genes. Long-awned plants are homozygous recessive, and awn-tipped plants homozygous dominant; glaucousness is dominant over non-glaucousness, and glabrous rachises over pubescent rachises. Awned plants appear to be established along the coasts of Newfoundland and Nova Scotia. They have generally been identified as Agropyron pungens (Pers.) Roem. & Schult., a species that has obtuse, mucronate lemmas.

Elymus repens is almost always a hexaploid. Most studies indicate that its genomic constitution is StStH, but Mason-Gamer (2001) demonstrated that it is genetically more complex than is implied by such a simple formula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 318. FNA vol. 24, p. 334.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Synonyms Elytrigia repens forma aristata, Elytrigia repens, Agropyron repens var. vaillantianum, Agropyron repens forma vaillantianum, Agropyron repens var. typicum, Agropyron repens forma trichorrhachis, Agropyron repens var. subulatum, Agropyron repens forma setiferum, Agropyron repens var. pilosum, Agropyron repens forma heberhachis, Agropyron repens forma aristatum, Agropyron repens
Name authority (J.G. Sm.) Burtt Davy (L.) Gould
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