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big squirreltail, big squirreltail grass

boreal wild rye, hairy wildrye, northern ryegrass, northwestern wildrye

Habit Plants cespitose, not rhizomatous. Plants cespitose, sometimes shortly rhizomatous.
Culms

15-65 cm, erect to ascending, usually puberulent;

nodes 4-6, mostly concealed, glabrous.

40-140 cm, usually somewhat decumbent;

nodes 4-7, mostly exposed, usually glabrous, occasionally puberulent.

Leaves

evenly distributed;

sheaths glabrous or white-villous;

auricles usually present, 0.5-1.5 mm;

ligules to 1 mm, truncate, entire or lacerate;

blades 1.5-4(5) mm wide, often ascending and involute, adaxial surfaces scabrous, pilose, or villous.

evenly distributed;

sheaths usually glabrous and smooth, occasionally scabridulous or retrorsely hairy, sometimes purplish;

auricles to 1.5 mm, often absent;

ligules to 1 mm;

blades 4-12 mm wide, lax, usually deep green, adaxial surfaces usually pilose or villous, occasionally puberulent or scabridulous.

Spikes

5-20 cm long, 5-15 cm wide, erect, sometimes partially enclosed at the base, with 2 spikelets per node, rarely with 3-4 at some nodes;

internodes 3-5(8) mm long, 0.1-0.3 mm thick at the thinnest sections, glabrous beneath the spikelets.

6-20 cm long, 0.5-2 cm wide, nodding to pendent, with 2 spikelets per node, rarely with 3 at some nodes;

internodes 3-10(12) mm long, 0.2-0.7 mm thick at the thinnest sections, usually glabrous, sometimes sparsely hairy.

Spikelets

10-15 mm, divergent, with 2-4 florets, lowest florets sterile and glumelike in 1 or both spikelets at each node;

disarticulation initially at the rachis nodes, subsequently beneath each floret.

12-20 mm, appressed to divergent, sometimes purplish at higher latitudes, with 2-4(7) florets, lowest florets functional;

disarticulation above the glumes and beneath each floret.

Glumes

subequal, (10)30-100 mm including the awns, the bases indurate and glabrous, glume bodies (2)5-10 mm long, 1-2 mm wide, setaceous, 2-3-veined, margins firm, awns (8)25-90 mm, each split into 3-9 unequal divisions, scabrous, flexuous to outcurving from near the glume bases at maturity;

fertile lemmas 8-10 mm, smooth or scabrous near the apices, 2 lateral veins extending into bristles to 10 mm, awns (10)20-110 mm long, about 0.2 mm wide at the base, divergent to arcuate;

paleas 7-9 mm, veins usually extending into about 1 mm bristles, apices acute to truncate;

anthers 1-2 mm.

equal or subequal, the bases flat, occasionally indurate for 0.5 mm, veins usually evident, glume bodies (4.5)7-10(11) mm long, 0.7-1.5 mm wide, linear-lanceolate, entire, widening or parallel-sided above the base, 3-5-veined, usually scabridulous to scabrous, veins occasionally hirsute beyond midlength, margins hyaline or scarious, awns 1-10 mm, straight;

lemmas 7-14 mm, smooth or scabridulous, lateral veins hairy, margins hairy beyond midlength, marginal hairs 0.5-1 mm, longer than those elsewhere, awns (2)8-30 mm, flexuous to moderately outcurving;

paleas 6-13 mm, with hairs of varying lengths on the keels and apices, acute, bidentate;

anthers 2-3.5 mm.

Anthesis

from late May to June.

from May to July.

2n

= 28.

= 28.

Elymus multisetus

Elymus hirsutus

Distribution
from FNA
AZ; CA; CO; ID; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; OR; WA; BC
[BONAP county map]
Discussion

Elymus multisetus grows in dry, often rocky, open woods and thickets on slopes and plains, from central Washington and Idaho to southern California, Colorado, and northwestern Arizona, and from sea level to 2000 m. It has also been reported from Baja California, Mexico. It usually grows in less arid habitats than E. elymoides subsp. elymoides (p. 319), but the two taxa are sometimes sympatric.

Wilson (1963) reported a wide belt of introgression between Elymus multisetus and E. elymoides subsp. elymoides from southeastern California to southern Nevada, but not in other areas where they are sympatric. There are also probable hybrids with E. glaucus (p. 306) and Pseudoroegneria spicata (p. 281).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Elymus hirsutus grows in moist to damp or dry soils in woods, thickets, and grasslands. Its range extends along the coastal mountains from the Aleutian Islands to northern Oregon, and inland to eastern British Columbia. Plants in the southern part of the range tend to have villous leaves and more erect spikes with shorter, straighter awns.

Elymus hirsutus is similar to E. glaucus (see previous), but its more pendent spikes, lemma pubescence pattern, and shorter glumes enable most specimens to be readily identified. Intermediates do exist; it is not known whether they reflect introgression or extremes of variation. Elymus hirsutus occasionally hybridizes with Leymus mollis (p. 356) and Hordeum brachyantherum (p. 243).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 318. FNA vol. 24, p. 310.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Elymus Poaceae > subfam. Pooideae > tribe Triticeae > Elymus
Sibling taxa
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hirsutus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
E. alaskanus, E. albicans, E. arizonicus, E. bakeri, E. canadensis, E. caninus, E. churchii, E. ciliaris, E. curvatus, E. dahuricus, E. diversiglumis, E. elymoides, E. glabriflorus, E. glaucus, E. hoffmannii, E. hystrix, E. interruptus, E. lanceolatus, E. macgregorii, E. macrourus, E. multisetus, E. pringlei, E. repens, E. riparius, E. scribneri, E. semicostatus, E. sibiricus, E. sierrae, E. stebbinsii, E. svensonii, E. texensis, E. trachycaulus, E. tsukushiensis, E. villosus, E. violaceus, E. virginicus, E. wawawaiensis, E. wiegandii, E. ×cayouetteorum, E. ×ebingeri, E. ×hansenii, E. ×palmerensis, E. ×pinalenoensis, E. ×pseudorepens, E. ×saundersii, E. ×yukonensis
Synonyms E. borealis
Name authority (J.G. Sm.) Burtt Davy J. Presl
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